Hoplocarida

Heterosquilla tricarinata, copyright Svamp.

Belongs within: Eumalacostraca.
Contains: Squillidae, Lysiosquillidae, Gonodactyloidea.

The Hoplocarida, including the mantis shrimps and their fossil relatives, are a group of malacostracan crustaceans bearing a small carapace and at least four free thoracic segments, together with a large abdomen ending in a tail fan.

<==Hoplocarida
    |  i. s.: Neogonodactylus RS10
    |           |--N. bredini SC17
    |           `--N. oerstedii RS10
    |--Perimecturidae [Palaeostomatopoda] BWW93
    |    |--Archaeocaris graffhami Brooks 1962 BWW93
    |    |--Eopteridium striatum Rohdendorf 1970 BWW93
    |    |--Bairdops beargulchensis Schram & Horner 1978 BWW93
    |    `--Perimecturus [incl. Eopterum; Archaeoptera] S77
    |         |--‘Eopterum’ devonicum Rohdendorf 1961 BWW93
    |         `--P. rapax Schram & Horner 1978 BWW93
    |--Aeschronectida G88
    |    |--Kallidechtes [Kallidecthidae] BWW93
    |    |    |--K. eageri Schram 1979 BWW93
    |    |    `--K. richardsoni Schram 1969 BWW93
    |    |--Aratidecthidae BWW93
    |    |    |--Crangopsis BWW93
    |    |    |    |--C. cooperi [=Pygocephalus cooperi] S74
    |    |    |    `--C. eskdalensis (Peach 1882) BWW93
    |    |    `--Aratidecthes johnsoni Schram 1969 BWW93
    |    `--Aenigmacarididae BWW93
    |         |--Joanellia BWW93
    |         |    |--J. elegans (Peach 1883) BWW93
    |         |    `--J. lundi G88
    |         `--Aenigmacaris G88
    |              |--A. cornigerum G88
    |              `--A. minima Schram & Schram 1979 BWW93
    `--Stomatopoda MD01
         |  i. s.: Alima hyalina Leach 1817 C73
         |         Pseudosquillopsis cerisii (Roux 1828) C73
         |         Eurysquilloides [Eurysquilloididae, Eurysquilloidoidea] MD01
         |         Erythrosquillidae [Erythrosquilloidea] MD01
         |--Tyrannophontidae [Archaeostomatopoda] BWW93
         |    |  i. s.: ‘Perimecturus’ pattoni Peach 1908 BWW93
         |    |--Tyrannophontes theridion BWW93
         |    `--Gorgonophontes peleron Schram 1984 BWW93
         `--Unipeltata BWW93
              |--Squillidae BWW93
              |--Bathysquilloidea BWW93
              |    |--Indosquillidae MD01
              |    |--Bathysquilla C73 [Bathysquillidae MD01]
              |    |    |--B. microps (Manning 1961) C73
              |    |    `--B. wetherelli (Woodward 1879) BWW93
              |    `--Sculdidae BWW93
              |         |--Pseudosculda laevis Schluter 1872 BWW93
              |         `--Sculda BWW93
              |              |--S. pennata Münster 1840 BWW93
              |              `--S. syriaca Dames 1886 BWW93
              |--Lysiosquilloidea BWW93
              |    |--Lysiosquillidae C73
              |    |--Coronis C73 [Coronididae MD01]
              |    |    `--C. excavatrix Brooks 1886 C73
              |    |--Nannosquilla C73 [Nannosquillidae MD01]
              |    |    |--N. occulta [=Lysiosquilla occulta] C73
              |    |    |--N. schmitti (Manning 1962) C73
              |    |    `--N. taylori Manning 1969 C73
              |    `--Heterosquilla K-MM02 [Heterosquillidae, Tetrasquillidae MD01]
              |         |--H. armata K-MM02
              |         |--H. brazieri HS01
              |         `--H. tricarinata HS01
              `--Gonodactyloidea BWW93

*Type species of generic name indicated

REFERENCES

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[C73] Camp, D. K. 1973. Stomatopod Crustacea. Memoirs of the Hourglass Cruises 3 (2): 1–100.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[K-MM02] Klein-MacPhee, G., & R. S. McBride. 2002. Sea robins. Family Triglidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3^rd ed. pp. 338–345. Smithsonian Institute Press: Washington.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[RS10] Regier, J. C., J. W. Shultz, A. Zwick, A. Hussey, B. Ball, R. Wetzer, J. W. Martin & C. W. Cunningham. 2010. Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences. Nature 463: 1079–1083.

[S74] Schram, F. R. 1974. Convergences between Late Paleozoic and modern caridoid Malacostraca. Systematic Zoology 23 (3): 323–332.

[S77] Schram, F. R. 1977. Paleozoogeography of Late Paleozoic and Triassic Malacostraca. Systematic Zoology 26 (4): 367–379.

[SC17] Schwentner, M., D. J. Combosch, J. P. Nelson & G. Giribet. 2017. A phylogenomic solution to the origin of insects by resolving crustacean-hexapod relationships. Current Biology 27: 1818–1824.

Astacidea

Enoplocytia leachii, from here.

Belongs within: Reptantia.
Contains: Astacida, Homarida.

The Astacidea are a clade of decapod crustaceans including the clawed lobsters and crayfish, with most modern representatives having well-developed chelae on the first pereiopods.

Characters (from Amati et al. 2004): Cephalothorax subcylindrical; rostrum and abdomen well developed; frontal portion of carapace not fused with epistome; antennae with five-segmented stalk and scale; third maxilliped pediform; pereiopods chelate, subchelate, or pseudochelate; first pereiopod longest; abdominal pleura well developed; exopods of uropods with diaresis; genital openings coxal.

Astacidea [Astacura, Nephropoidea]
    |  i. s.: Platychelidae BWW93
    |           |--Platypleon nevadense Van Straelen 1936 BWW93
    |           `--Platychela trauthi Glaessner 1931 BWW93
    |         Viriliastacus AFZ04
    |--+--+--Astacida AFZ04
    |  |  `--Homarida DAS03
    |  `--Thaumastochelida DAS03
    |       |--Thaumastochelopsis Bruce 1988 TS03
    |       |--Oncopareia Bosquet 1854 TS03
    |       |    `--O. bredai TS03
    |       `--Thaumastocheles Wood-Mason 1874 [Thaumastochelidae] TS03
    |            |--T. dochmiodon Chan & de Saint Laurent 1999 DAS03
    |            `--T. japonicus TS03
    `--+--Glypheoidea AFZ04
       |    |--Chimaerastacus Amati, Feldmann & Zonneveld 2004 [Chimaerastacidae] AFZ04
       |    |    `--*C. pacifluvialis Amati, Feldmann & Zonneveld 2004 AFZ04
       |    `--+--Neoglyphea inopinata Forest & de Saint Laurent 1975 AFZ04, DAS03
       |       |--Glypheidae FGS04
       |       |    |--Glyphea robusta AFZ04, P98
       |       |    |--Protoclytiopsis antiqua Birshtein 1958 BWW93
       |       |    `--Litogaster von Meyer 1847 [=Lithogaster (l. c.)] FGS04
       |       |         `--L. turnbullensis AFZ04
       |       `--Mecochiridae AFZ04
       |            |  i. s.: ‘Upogebia’ clypeata BWW93
       |            |--Mecochirus AFZ04
       |            |--Meyeria M’Coy 1849 BWW93
       |            `--Pseudoglyphea Oppel 1861 BWW93 [incl. Triasiglyphea van Straelen 1936 AFZ04]
       |                 `--P. mulleri (van Straelen 1936) [=*Triasiglyphea mulleri] AFZ04
       `--Erymoidea AFZ04
            |--Clytiopsis [Clytiopsidae] AFZ04
            |    `--C. argentoratensis Glaessner 1929 BWW93
            `--+--Pemphicidae AFZ04
               |    |--Pseudopemphix Wüst 1903 BWW93
               |    |    `--P. albertii (von Meyer 1840) BWW93
               |    `--Pemphix von Meyer 1840 FGS04
               |         `--P. sueurii (Desmarest 1822) BWW93
               `--Erymidae AFZ04
                    |  i. s.: Lissocardia von Meyer 1851 FGS04
                    |--Eryma von Meyer 1840 AFZ04, TS03
                    `--+--Phlyctisoma AFZ04
                       `--Enoplocytia McCoy 1849 AFZ04, E14
                            |--*E. leachii [=Astacus leachii] E14
                            |--E. terrareginae Etheridge 1914 E14
                            `--E. ventricosa E14

*Type species of generic name indicated

REFERENCES

[AFZ04] Amati, L., R. M. Feldmann & J.-P. Zonneveld. 2004. A new family of Triassic lobsters (Decapoda: Astacidea) from British Columbia and its phylogenetic context. Journal of Paleontology 78 (1): 150–168.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[DAS03] Dixon, C. J., S. T. Ahyong & F. R. Schram. 2003. A new hypothesis of decapod phylogeny. Crustaceana 76: 935–975.

[E14] Etheridge, R., Jr. 1914. The genus Enoploclytia in the Cretaceous rocks of Queensland. Records of the Australian Museum 10 (9): 271–273, pls 23–24.

[FGS04] Feldmann, R. M., A. A. Garassino & C. E. Schweitzer. 2004. The presumed decapod, Palaeopemphix Gemmellaro, 1890, is a unique member of the Phyllocarida (Palaeopemphicida: Palaeopemphidae). Journal of Palaeontology 78: 340–348.

[P98] Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.

[TS03] Tshudy, D., & U. Sorhannus. 2003. Hoploparia, the best-known fossil clawed lobster (family Nephropidae), is a “wastebasket” genus. Journal of Crustacean Biology 23 (3): 700–711.

Potamoidea

Potamon fluviatilis, copyright Fabio Liverani.

Belongs within: Gecarcinucidae.
Contains: Potamonautidae.

The Potamoidea are a group of freshwater and semi-terrestrial crabs found in tropical and subtropical regions of the Old World. Members are characterised by males having the distal section of the second gonopods forming a closed tube.

See also: Crabs in rivers, crabs in trees.

Potamoidea
    |--Sundathelphusidae H86
    |    |--Perithelphusa borneensis (von Martens 1868) H86
    |    |--Sundathelphusa tenebrosa Holthuis 1979 H86
    |    `--Holthuisana Bott 1969 H86
    |         `--H. alba Holthuis 1980 H86
    `--Potamidae [Sinopotamidae, Thelphusidae] BS05
         |  i. s.: Cerberusa Holthuis 1979 H86
         |           |--C. coeca Holthuis 1979 H86
         |           `--C. tipula Holthuis 1979 H86
         |         Paratelphusula Alcock 1909 (see below for synonymy) BS05
         |           |--*P. dayana (Wood-Mason 1871) (see below for synonymy) BS05
         |           |--P. calvum (Alcock 1909) [=Potamon (Paratelphusula) calvum] BS05
         |           |--P. crenulifera (Wood-Mason 1871) (see below for synonymy) BS05
         |           |--P. feae (de Man 1898) [=Parathelphusa feae] BS05
         |           |--P. fungosum (Alcock 1909) [=Potamon (Paratelphusula) fungosum] BS05
         |           |--P. martensi (Wood-Mason 1875) (see below for synonymy) BS05
         |           |--P. panningi (Bott 1966) [=Potamon (Spinopotamon) panningi, Acanthopotamon panningi] BS05
         |           `--P. woodmasoni (Rathbun 1905) [=Potamon (Parathelphusa) woodmasoni] BS05
         |         Himalayapotamon Pretzmann 1966 BS05
         |           |--*H. atkinsoniana (Wood-Mason 1871) (see below for synonymy) BS05
         |           |--H. emphysetum (Alcock 1910) (see below for synonymy) BS05
         |           `--H. sunkoshiense Brandis & Sharma 2005 BS05
         |         Potamiscus Alcock 1909 [incl. Ranguna Bott 1966, Terrapotamon Ng 1986] BS05
         |           |--*P. annandalii Alcock 1909 [=Potamon (*Potamiscus) annandalii] BS05
         |           |--‘Potamon’ abbotti Rathbun 1898 [=*Terrapotamon abbotti] BS05
         |           |--P. koolooense (Rathbun 1904) [incl. Potamon babaulti Bouvier 1918] BS05
         |           |--P. rangoonensis Bott 1966 [=*Ranguna rangoonensis] BS05
         |           `--P. sikkimensis (Rathbun 1905) (see below for synonymy) BS05
         |         Parapotamon de Man 1907 BS05
         |           `--*P. spinescens (Calman 1905) BS05
         |         Stoliczia BS05
         |         Archithelphusa punctulatus (Heer 1865) BWW93
         |         Pseudopotamon speciosum (Capellini 1874) BWW93
         |--+--Socotra pseudocardisoma DC06
         |  `--+--Johora tiomanensis DC06
         |     `--Geothelphusa DC06
         |          |--G. albogilva DC06
         |          `--G. dehaanii (White 1847) H86
         `--+--Potamonautidae DC06
            `--+--Isolapotamon [Isolapotamidae] DC06
               |    |--I. collinsi Holthuis 1979 H86
               |    `--I. consobrinum DC06
               `--Potamon Savigny 1816 DC06, BS05 [incl. Thelphusa Latreille 1819 BS05]
                    |--P. fluviatilis DC06
                    |    |--P. f. fluviatilis H86
                    |    `--P. f. algeriense Bott 1967 H86
                    |--P. rangoonense Rathbun 1904 BS05
                    `--P. sivaleuse G88

*Himalayapotamon atkinsoniana (Wood-Mason 1871) [=Telphusa atkinsoniana, Potamon (*Himalayapotamon) atkinsonianum; incl. P. (H.) atkinsonianum janetschekii Pretzmann 1966] BS05

Himalayapotamon emphysetum (Alcock 1910) [=Potamon atkinsonianum var. emphysetum; incl. P. (Himalayapotamon) atkinsonianum gordoni Pretzmann 1966, P. atkinsonianum var. ventriosum Alcock 1910] BS05

Paratelphusula Alcock 1909 [incl. Acanthopotamon Kemp 1918, Lobothelphusa Bouvier 1917, Spinopotamon Bott 1966] BS05

Paratelphusula crenulifera (Wood-Mason 1871) [=Paratelphusa crenulifera, Hydrothelphusa (*Lobothelphusa) crenulifera] BS05

*Paratelphusula dayana (Wood-Mason 1871) [=Paratelphusa dayana, Potamon (*Paratelphusula) dayanum, Telphusa (Parathelphusa) dayana] BS05

Paratelphusula martensi (Wood-Mason 1875) [=Paratelphusa martensi, *Acanthopotamon martensi, Potamon (Acanthotelphusa) martensi, Po. (Parathelphusa) martensi, Po. (*Spinopotamon) martensi] BS05

Potamiscus sikkimensis (Rathbun 1905) [=Potamon (Geothelphusa) sikkimensis, Potamon (Potamiscus) sikkimense; incl. Potamon tumidulum Alcock 1909] BS05

*Type species of generic name indicated

REFERENCES

[BS05] Brandis, D., & S. Sharma. 2005. Taxonomic revision of the freshwater crab fauna of Nepal with description of a new species (Crustacea, Decapoda, Brachyura, Potamoidea and Gecarcinucoidea). Senckenbergiana Biologica 85: 1–30.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[DC06] Daniels, S. R., N. Cumberlidge, M. Pérez-Losada, S. A. E. Marijnissen & K. A. Crandall. 2006. Evolution of Afrotropical freshwater crab lineages obscured by morphological convergence. Molecular Phylogenetics and Evolution 40 (1): 227–235.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr W. Backhuys: Leiden.

Procambarus

Red swamp crayfish Procambarus clarkii, copyright Greg Peterson.

Belongs within: Astacida.

Procambarus is a North American genus of fairly generalised freshwater crayfish.

<==Procambarus Ortmann 1905 H86
    |--P. (Austrocambarus Hobbs 1972) H86
    |    |--P. (A.) cubensis H86
    |    |    |--P. c. cubensis H86
    |    |    `--P. c. rivalis (Faxon 1912) H86
    |    |--P. (A.) mirandai Villalobos 1954 H86
    |    |--P. (A.) niveus Hobbs & Villalobos 1964 H86
    |    |--P. (A.) oaxacae Hobbs 1973 H86
    |    |    |--P. o. oaxacae H86
    |    |    `--P. o. reddelli Hobbs 1973 H86
    |    |--P. (A.) rodriguezi Hobbs 1943 H86
    |    `--P. (A.) sbordonii Hobbs 1977 H86
    |--P. (Girardiella) simulans (Faxon 1884) H86
    |--P. (Leconticambarus Hobbs 1972) H86
    |    |--P. (L.) barbatus (Faxon 1890) [=Cambarus barbatus] H86
    |    `--P. (L.) milleri Hobbs 1971 H86
    |--P. (Lonnbergius Hobbs 1972) H86
    |    `--P. (L.) acherontis (Lönnberg 1894) H86
    |--P. (Ortmannicus Fowler 1912) H86
    |    |--P. (O.) acutus (Girard 1952) H86
    |    |    |--P. a. acutus H86
    |    |    `--P. a. cuevachicae (Hobbs 1941) H86
    |    |--P. (O.) erythrops Relyea & Sutton 1975 H86
    |    |--P. (O.) fallax (Hagen 1870) H86
    |    |--P. (O.) franzi Hobbs & Lee 1976 H86
    |    |--P. (O.) horsti Hobbs & Means 1972 H86
    |    |--P. (O.) leitheuseri Franz & Hobbs 1983 H86
    |    |--P. (O.) lucifugus (Hobbs 1940) H86
    |    |    |--P. l. lucifugus H86
    |    |    `--P. l. alachua (Hobbs 1940) H86
    |    |--P. (O.) orcinus Hobbs & Means 1972 H86
    |    |--P. (O.) pallidus (Hobbs 1940) H86
    |    |--P. (O.) toltecae (Hobbs 1943) H86
    |    `--P. (O.) villalobosi Hobbs 1969 H86
    |--P. (Remoticambarus Hobbs 1972) H86
    |    `--P. (R.) pecki Hobbs 1967 H86
    |--P. (Scapulicambarus Hobbs 1972) H86
    |    |--P. (S.) clarkii (Girard 1852) H86
    |    |--P. (S.) paeninsulanus (Faxon 1914) H86
    |    `--P. (S.) xilitlae Hobbs & Grubbs 1982 H86
    `--P. (Villalobosus) H86
         |--P. (V.) cuetzalanae Hobbs 1982 H86
         `--P. (V.) xochitlanae Hobbs 1975 H86

Procambarus incertae sedis:
  P. leonensis GR98
  Spiculifer group H56
    |--P. dupratzi Penn 1953 H56
    |--P. echinatus Hobbs 1956 H56
    |--P. natchitochae Penn 1953 H56
    |--P. penni Hobbs 1951 H56
    |--P. raneyi Hobbs 1953 H56
    |--P. spiculifer (LeConte 1856) H56
    |--P. suttkusi Hobbs 1953 H56
    |--P. versutus (Hagen 1870) H56
    `--P. vioscai Penn 1946 H56

*Type species of generic name indicated

REFERENCES

[GR98] Giribet, G., & C. Ribera. 1998. The position of arthropods in the animal kingdom: a search for a reliable outgroup for internal arthropod phylogeny. Molecular Phylogenetics and Evolution 9: 481–488.

[H56] Hobbs, H. H., Jr. 1956. A new crayfish of the genus Procambarus from South Carolina (Decapoda: Astacidae). Journal of the Washington Academy of Sciences 46 (4): 117–121.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr W. Backhuys: Leiden.

Syncarida

Allobathynella japonica, from Morimoto & Miura (1957).

Belongs within: Eumalacostraca.
Contains: Bathynella, Hexabathynella, Iberobathynella.

The Syncarida are a group of malacostracan crustaceans found primarily in fresh water that lack a carapace. Living examples are divided between the Anaspidacea, in which the first thoracic segment is fused to the head and a tail fan is present, and the minute Bathynellacea that lack a tail fan.

<==Syncarida S86
    |  i. s.: Allanaspides S86
    |--Palaeocaridacea MD01
    |    |--Clarkecaris [Clarkecarididae] G88
    |    |    `--C. brazilicus (Clarke 1920) S77
    |    |--Minicarididae BWW93
    |    |    |--Minicaris brandi Schram 1979 BWW93
    |    |    `--Erythrogaulos carrizoensis Schram 1984 BWW93
    |    |--Squillitidae BWW93
    |    |    |--Squillites spinosa Scott 1938 BWW93
    |    |    `--Nectotelson krejcii (Fritsch 1875) BWW93
    |    |--Palaeocaris [Palaeocarididae] G88
    |    |    |--P. canadensis G88
    |    |    |--P. retractata Calman 1932 BWW93
    |    |    |--P. secretanae Schram 1984 BWW93
    |    |    `--P. typus G88
    |    `--Acanthotelsonidae BWW93
    |         |--Palaeosyncaris dakotensis Brooks 1962 BWW93
    |         |--Acanthotelson [incl. Eileticus] WA04
    |         |    |--‘Eileticus’ anthracinus Scudder 1882 WA04
    |         |    `--A. stimpsoni G88
    |         `--Uronectes BWW93
    |              |--U. fimbriatus (Jordan 1947) BWW93
    |              `--U. kinniensis G88
    |--Anaspidacea [Anaspidinea] LSE13
    |    |--Anaspididae BWW93
    |    |    |--Anaspidites antiquus (Chilton 1929) BWW93
    |    |    |--Anaspides Thomson 1894
    |    |    |    `--A. tasmaniae (Thomson 1893)
    |    |    `--Koonaspides indistinctus G88
    |    |--Koonungidae S86
    |    |    |--Koonunga N31
    |    |    `--Micraspides Nicholls 1931 N31
    |    |         `--*M. calmani Nicholls 1931 N31
    |    |--Psammaspididae S86
    |    |    |--Psammaspides Schminke 1974 S86
    |    |    |    `--P. williamsi S86
    |    |    `--Eucrenonaspides Knott & Lake 1980 S86
    |    |         `--E. oinotheke Knott & Lake 1980 S86
    |    `--Stygocarididae [Stygocaridacea] S86
    |         |--Oncostygocaris Schminke 1980 S86
    |         |    `--O. patagonica (Noodt 1963) S86
    |         |--Stygocarella Schminke 1980 S86
    |         |    `--S. pleotelson Schminke 1980 S86
    |         |--Parastygocaris Noodt 1963 S86
    |         |    |--P. andina Noodt 1963 S86
    |         |    `--P. goerssi Noodt 1963 S86
    |         `--Stygocaris Noodt 1963 S86
    |              |--S. giselae Schminke 1980 S86
    |              |--S. gomezmillasi Noodt 1963 S86
    |              `--S. townsendi Morimoto 1977 S86
    `--Bathynellacea LSE13
         |--Bathynellidae S86
         |    |--Bathynella S86
         |    |--Sardobathynella Serban 1973 S86
         |    |    `--S. cottarelli Serban 1973 S86
         |    |--Vandelibathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    `--V. vandeli (Delamare Deboutteville & Chappuis 1954) S86
         |    |--Agnathobathynella Schminke 1980 S86
         |    |    `--A. ecclesi Schminke 1980 S86
         |    |--Austrobathynella Delamare Deboutteville 1960 S86
         |    |    `--A. patagonica Delamare Deboutteville 1960 S86
         |    |--Transvaalthynella Serban & Coineau 1975 S86
         |    |    `--T. coineaui Serban & Coineau 1975 S86
         |    |--Transkeithynella Serban & Coineau 1975 S86
         |    |    `--T. paradoxa Serban & Coineau 1975 S86
         |    |--Parameridiobathynella Serban & Leclerc 1984 S86
         |    |    `--P. gardensis Serban & Leclerc 1984 S86
         |    |--Pseudobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    |--P. helvetica Serban & Delamare Deboutteville 1974 S86
         |    |    `--P. magniezi Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |--Meridiobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    |--M. catalanensis Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    `--M. rouchi Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |--Vejdovskybathynella Serban & Leclerc 1984 S86
         |    |    |--V. balazuci Serban & Leclerc 1984 S86
         |    |    `--V. espattyensis Serban & Leclerc 1984 S86
         |    |--Antrobathynella Serban 1966 S86
         |    |    `--A. stammeri (Jakobi 1954) S86
         |    |         |--A. s. stammeri S86
         |    |         `--A. s. ciscaucasia (Birstein & Ljovuschkin 1964) S86
         |    |--Nannobathynella Noodt 1969 S86
         |    |    |--N. africana Schminke & Wells 1974 S86
         |    |    |--N. eburnea Schminke 1979 S86
         |    |    `--N. marcusi Noodt 1969 S86
         |    `--Gallobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. boui Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. coiffaiti (Delamare Deboutteville 1961) S86
         |         |--G. delayi Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. juberthiae Serban, Coineau & Delamare Deboutteville 1971 S86
         |         `--G. tarissei Serban, Coineau & Delamare Deboutteville 1971 S86
         `--Parabathynellidae S86
              |--Lamtobathynella pentodonta S86
              |--Hexabathynella S86
              |--Iberobathynella S86
              |--Batubathynella Schminke 1973 S86
              |    `--B. malaya (Sars 1929) S86
              |--Nipponbathynella Schminke 1973 S86
              |    `--N. miurai (Ueno 1952) S86
              |--Ctenophallonella Coineau & Serban 1978 S86
              |    `--C. mutlumuviensis Coineau & Serban 1978 S86
              |--Haplophallonella Serban & Coineau 1975 S86
              |    `--H. heterodonta Serban & Coineau 1975 S86
              |--Heterodontobathynella Schminke 1973 S86
              |    `--H. ninianae (Green 1964) S86
              |--Brasilibathynella Jakobi 1958 S86
              |    `--B. florianopolis Jakobi 1958 S86
              |--Odontobathynella Delamare Deboutteville & Serban 1979 S86
              |    `--O. amazonica Delamare Deboutteville & Serban 1979 S86
              |--Acanthobathynella Coineau 1967 S86
              |    `--A. knoepffleri Coineau 1967 S86
              |--Afrobathynella Schminke 1976 S86
              |    `--A. trimera Schminke 1976 S86
              |--Nunubathynella Schminke 1976 S86
              |    `--N. dimera Schminke 1976 S86
              |--Psalidobathynella Schminke 1979 S86
              |    `--P. stocki Schminke 1979 S86
              |--Nilobathynella Dumont 1984 S86
              |    `--N. predynastica Dumont 1984 S86
              |--Thermobathynella Capart 1951 S86
              |    |--T. adami Capart 1951 S86
              |    `--T. amyxi Siewing 1956 S86
              |--Habrobathynella Schminke 1973 S86
              |    |--H. jeanneli (Delamare Deboutteville & Paulian 1954) S86
              |    `--H. milloti (Delamare Deboutteville & Paulian 1954) S86
              |--Chilibathynella Noodt 1963 S86
              |    |--C. australiensis Schminke 1973 S86
              |    `--C. clandestina Noodt 1963 S86
              |--Noodtibathynella Schminke 1973 S86
              |    |--N. jumboli (Siewing 1958) S86
              |    `--N. neotropica (Noodt 1965) S86
              |--Parabathynella Chappuis 1926 S86
              |    |--P. malaya N31
              |    |--P. motasi Dancau & Serban 1963 S86
              |    `--P. stygia Chappuis 1926 [incl. P. phreatica Chappuis 1939] S86
              |--Parvulobathynella Schminke 1973 S86
              |    |--P. camposicola (Jakobi 1961) S86
              |    |--P. riegelorum (Noodt 1965) S86
              |    `--P. ypacaraiensis (Noodt 1963) S86
              |--Leptobathynella Noodt 1965 [Leptobathynellidae] S86
              |    |--L. camposicola (Jakobi 1961) S86
              |    `--L. richerti (Noodt 1963) S86
              |         |--L. r. richerti S86
              |         `--L. r. brasiliensis Noodt 1972 S86
              |--Atopobathynella Schminke 1973 S86
              |    |--A. chelifera Schminke 1973 S86
              |    |--A. compagana Schminke 1973 S86
              |    |--A. hospitalis Schminke 1973 S86
              |    `--A. valdiviana (Noodt 1965) S86
              |--Eobathynella Birstein & Ljovuschkin 1964 S86
              |    |--E. gracillima (Ueno 1956) S86
              |    |--E. matuta (Morimoto 1970) S86
              |    |--E. mesasiatica (Birstein & Ljovuschkin 1964) S86
              |    |--E. minima Jankowskaja 1972 S86
              |    `--E. tianshanica (Jankowskaja 1964) S86
              |--Notobathynella Schminke 1973 S86
              |    |--N. chiltoni Schminke 1973 S86
              |    |--N. hineoneae Schminke 1973 S86
              |    |--N. longipes Schminke 1978 S86
              |    |--N. remota Schminke 1973 S86
              |    `--N. williamsi Schminke 1973 S86
              |--Cteniobathynella Schminke 1973 S86
              |    |--C. bakeri (Green 1964) S86
              |    |--C. calmani (Por 1968) S86
              |    |--C. caparti (Fryer 1957) S86
              |    |--C. essameuri Dumont 1981 S86
              |    |--C. leleupi (Delamare Deboutteville & Chappuis 1955) S86
              |    |--C. noodti Schminke 1973 S86
              |    `--C. teocchii (Coineau & Knoepffler 1971) S86
              `--Allobathynella Morimoto & Miura 1957 S86
                   |--A. carinata (Ueno 1952) S86
                   |--A. coreana Morimoto 1970 S86
                   |--A. gigantea (Morimoto 1959) S86
                   |    |--A. g. gigantea S86
                   |    `--A. g. pluto (Morimoto 1963) S86
                   |--A. japonica Morimoto & Miura 1957 S86
                   |--A. kuma (Ueno 1956) S86
                   |--A. mirabilis Ueno 1961 S86
                   `--A. yaye (Ueno 1956) S86

*Type species of generic name indicated

REFERENCES

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[LSE13] Legg, D. A., M. D. Sutton & G. D. Edgecombe. 2013. Arthropod fossil data increase congruence of morphological and molecular phylogenies. Nature Communications 4: 2485.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[N31] Nicholls, G. E. 1931. Micraspides calmani, a new syncaridan from the west coast of Tasmania. Linnean Society's Journal—Zoology 37 (254): 473–488.

[S86] Schminke, H. K. 1986. Syncarida. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 389–404. E. J. Brill/Dr W. Backhuys: Leiden.

[S77] Schram, F. R. 1977. Paleozoogeography of Late Paleozoic and Triassic Malacostraca. Systematic Zoology 26 (4): 367–379.

[WA04] Wilson, H. M., & L. I. Anderson. 2004. Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland. Journal of Paleontology 78 (1): 169–184.

Haustoroidea

Eohaustorius washingtonianus, copyright Florida Museum of Natural History.

Belongs within: Amphipoda.

The Haustoroidea are a Holarctic group of marine amphipods with a short, broad, weakly rostrate head, 'pseudorostrate' paired peduncles on antenna 1, weakly or non-dactylate peraeopods and maxilliped, and a strongly deflexed urosome (Bousfield & Hoover 1995).

<==Haustoroidea [Pontoporeioidea] R99
    |--Cheidae R99
    |--Condukiidae R99
    |--Ipanemidae R99
    |--Sinurothoe Ren 1999 [Sinurothoidae] R99
    |    `--*S. sinensis Ren 1999 R99
    |--Cardenio Stebbing 1888 B69 [Cardenioidae R99]
    |    `--*C. paurodactylus Stebbing 1888 B69
    |--Platyischnopus Stebbing 1888 B69 [Platyischnopidae R99]
    |    |--*P. mirabilis Stebbing 1888 B69
    |    `--P. gracilipes Schellenberg 1931 [incl. Phoxocephalus capuciatus Oliveira 1955] B69
    |--Pontoporeiidae R99
    |    |--Monoporeia affinis BH95, VVP01 [=Pontoporeia affinis VVP01]
    |    |--Diporeia BH95
    |    |--Priscillina Stebbing 1888 BH95, B69 [=Priscilla Boeck 1871 (preoc.) B69]
    |    |    `--*P. armata (Boeck 1861) [=Pontoporeia armata, *Priscilla armata] B69
    |    `--Pontoporeia Krøyer 1842 B69
    |         `--*P. femorata Krøyer 1842 B69 [incl. P. furcigera Bruzelius 1859 N00]
    |--Urothoe Dana 1852 B69 [incl. Egidia Costa 1853 B69; Urothoidae R99]
    |    |--*U. irrostratus Dana 1853 B69
    |    |--U. abbreviata N00
    |    |--U. brevicornis Bate 1862 BD95
    |    |--U. cuspis R99
    |    |--U. elegans Bate 1857 [incl. U. norvegica Boeck 1861] B69
    |    |--U. grimaldii Chevreux 1895 BD95
    |    |--U. intermedia Bellan-Santini & Ruffo 1986 BD95
    |    |--U. marina (Bate 1857) BD95 [=U. marinus (l. c.) BK77; incl. U. bairdii Bate & Westwood 1862 N00]
    |    |--U. norvegica Boeck 1876 N00
    |    |--U. poseidonis Reibisch 1905 BD95
    |    |--U. poucheti N00
    |    `--U. pulchella (Costa 1853) BD95 [=Egidia pulchella N00]
    `--Haustoriidae BH95
         |--Pseudohaustorius Bousfield 1965 B69
         |    `--*P. caroliniensis Bousfield 1965 B69
         |--Protohaustorius Bousfield 1965 B69
         |    `--*P. deichmannae Bousfield 1965 B69
         |--Acanthohaustorius Bousfield 1965 B69
         |    `--*A. millsi Bousfield 1965 B69
         |--Neohaustorius Bousfield 1965 B69
         |    `--*N. schmitzi Bousfield 1965 B69
         |--Parahaustorius Bousfield 1965 B69
         |    `--*P. longimerus Bousfield 1965 B69
         |--Urothoides Stebbing 1891 B69
         |    `--*U. lachneessa (Stebbing 1888) [=Urothoe lachneessa] B69
         |--Haustorius Můller 1775 (see below for synonymy) B69
         |    |--*H. arenarius (Slabber 1769) B69 (see below for synonymy)
         |    `--H. algeriensis Mulot 1968 BD95
         |--Carangolia Barnard 1961 B69
         |    |--*C. mandibularis Barnard 1962 B69
         |    `--C. puliciformis Barnard 1961 B69
         `--Eohaustorius Barnard 1957 BH95
              |--*E. washingtonianus (Thorsteinson 1941) [=Haustorius washingtonianus] BH95
              |--E. barnardi Bousfield & Hoover 1995 BH95
              |--E. brevicuspis Bosworth 1973 BH95
              |--E. cheliferus (Bulycheva 1952) [=Haustorius cheliferus, E. heliferus (l. c.)] BH95
              |--E. eous (Gurjanova 1951) [=Haustorius eous] BH95
              |--E. estuarius Bosworth 1973 BH95
              |--E. gurjanovae Bousfield & Hoover 1995 BH95
              |--E. longicarpus Bousfield & Hoover 1995 BH95
              |--E. robustus (Gurjanova 1953) [=Haustorius eous robustus] BH95
              |--E. sawyeri Bosworth 1973 BH95
              |--E. sencillus Barnard 1962 BH95
              |--E. subulicolus Hirayama 1985 [=E. subulicola] BH95
              `--E. tandeensis Dang 1968 BH95

Haustorius Můller 1775 [incl. Bellia Bate 1851 (preoc.), Lepidactylis Say 1818, Pterygocerus Latreille 1825, Sulcator Bate 1854] B69

*Haustorius arenarius (Slabber 1769) B69 [=Oniscus arenarius B69, Gammarus arenarius L02, Pterygocera arenaria N00, Sulcator arenarius N00; incl. Lepidactylis dytiscus Say 1818 N00]

*Type species of generic name indicated

REFERENCES

[BD95] Bakalem, A., & J.-C. Dauvin. 1995. Inventaire des crustacés amphipodes (Gammaridea, Caprellidea, Hyperiidea) des costes d'Algérie: essai de synthèse. Mésogée 54: 49–62.

[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1–159.

[B69] Barnard, J. L. 1969. The families and genera of marine gammaridean Amphipoda. United States National Museum Bulletin 271: i–vi, 1–535.

[BH95] Bousfield, E. L., & P. Hoover. 1995. The amphipod superfamily Pontoporeioidea on the Pacific coast of North America. I. Family Haustoriidae. Genus Eohaustorius J. L. Barnard: systematics and distributional ecology. Amphipacifica 2: 35–63.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[N00] Norman, A. M. 1900. British Amphipoda: families Pontoporeidae to Ampeliscidae. Annals and Magazine of Natural History, series 7, 5: 326–346.

[R99] Ren X.-Q. 1999. A new family of superfamily Haustoroidea (Crustacea: Amphipoda: Gammaridea) from the China Sea. Chinese Journal of Oceanology & Limnology 17: 344–349.

[VVP01] Väinölä, R., J. K. Vainio & J. U. Palo. 2001. Phylogeography of “glacial relict” Gammaracanthus (Crustacea, Amphipoda) from boreal lakes and the Caspian and White Seas. Canadian Journal of Fisheries and Aquatic Science 58: 2247–2257.

Palaemonetes

Palaemonetes pugio, copyright Brian Gratwicke.

Belongs within: Palaemonidae.

Palaemonetes, glass shrimp, is a cosmopolitan genus of freshwater shrimp that typically lack a mandibular palp.

<==Palaemonetes Heller 1869 H86
    |  i. s.: P. antennarius (Milne Edwards 1837) H86
    |         P. argentinus CB93
    |         P. exilipes M85
    |         P. kadiakensis M85
    |         P. varians CS77
    |--P. (Palaemonetes) H86
    |    |--P. (P.) cummingi Chace 1954 H86
    |    |--P. (P.) intermedius M85
    |    |--P. (P.) paludosus (Gibbes 1850) M85, H86
    |    |--P. (P.) pugio Holthuis 1949 M85, M88
    |    `--P. (P.) vulgaris (Say 1818) M85, M88
    `--P. (Alaocaris Holthuis 1949) H86
         |--P. (A.) antrorum Benedict 1896 H86
         `--P. (A.) holthuisi Strenth 1976 H86

*Type species of generic name indicated

REFERENCES

[CB93] Cash, C. E., & R. T. Bauer. 1993. Adaptations of the branchial ectoparasite Probopyrus pandalicola (Isopoda: Bopyridae) for survival and reproduction related to ecdysis of the host, Palaemonetes pugio (Caridea: Palaemonidae). Journal of Crustacean Biology 13 (1): 111–124.

[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr W. Backhuys: Leiden.

[M85] Markham, J. C. 1985. A review of the bopyrid isopods infesting caridean shrimps in the northwestern Atlantic Ocean, with special reference to those collected during the Hourglass Cruises in the Gulf of Mexico. Memoirs of the Hourglass Cruises 7 (3): 1–156.

[M88] Markham, J. C. 1988. Descriptions and revisions of some species of Isopoda Bopyridae of the northwestern Atlantic Ocean. Zoologische Verhandelingen 246: 1–63.

Lysianassidae

Orchomene amblyops, copyright University of Bergen.

Belongs within: Amphipoda.

The Lysianassidae are a group of marine amphipods characterised by a mitten-like terminal section on the section gnathopod (Barnard 1969).

Characters (from Barnard 1969): Accessory flagellum usually present and more than 3-articulate but occasionally vestigial or few in segments; peduncle of antenna 1 short and stout, articles 2 and 3 usually much shorter than 1 and occasionally scarcely distinguishable because of telescoping; pereon generally lacking processes, often urosomal segment 1 processiferous; body compact, chitin usually very smooth, porcellanous; mouthparts enormously variable, either chewing or piercing, often with reduction in mandibular molars, variable in position of mandibular palps; upper lip and epistome invaluable for identification, either or both often produced in variable configurations; maxilla 1 generally with 2-articulate palp but occasionally reduced or absent; maxilla 2 occasionally poorly developed; maxilliped occasionally with reduced plates and palps; gnathopods usually quite small, 2 mitten-shaped terminally, with article 3 elongate, article 5 generally with a "pineapple cushion", i.e., posterior margin bulbous and with microscopic texture of a pineapple, structure composed of scales or coarse setules, article 6 generally covered with coarse setules or tasseled setules; pereopods generally stout, occasionally prehensile; coxae usually long, first one or two or three occasionally reduced in size or hidden by following coxae; rami of uropod 3 lanceolate, outer often 2-articulate, inner occasionally reduced in size, rarely both absent; telson cleft, emarginate or entire, long or shorty rarely absent.

<==Lysianassidae
    |--Lucayarina catacumba Clark & Barnard 1985 S86
    |--Acontiostoma Stebbing 1888 B69
    |    `--*A. marionis Stebbing 1888 B69 [incl. A. magellanicum N38]
    |--Paralysianopsis Schellenberg 1931 [incl. Austronisimus Barnard 1931] B69
    |    `--*P. odhneri Schellenberg 1931 B69
    |--Charcotia Chevreux 1905 [=Waldeckia Chevreux 1906; incl. Ephippiphora White 1847 (preoc.)] B69
    |    `--*C. obesa Chevreux 1905 B69 [=*Waldeckia obesa B69; incl. W. zshauii non Anonyx zschauii N38]
    |--Pseudorchomene Schellenberg 1926 B69
    |    `--*P. coatsi (Chilton 1912) B69 [=Orchomenopsis coatsi N38]
    |--Aroui Chevreux 1911 B69
    |    `--*A. setosus Chevreux 1911 B69
    |--Perrierella Chevreux & Bouvier 1892 [incl. Pararistias Robertson 1892] B69
    |    `--*P. audouiniana (Bate 1857) B69 (see below for synonymy)
    |--Socarnopsis Chevreux 1911 B69
    |    `--*S. crenulata Chevreux 1911 B69
    |--Adeliella Nicholls 1938 N38
    |    `--*A. laticornis Nicholls 1938 N38
    |--Alicella Chevreux 1899 B69
    |    `--*A. gigantea Chevreux 1899 B69
    |--Allogaussia Schellenberg 1926 B69
    |    `--*A. paradoxa Schellenberg 1926 B69
    |--Amaryllis Haswell 1880 non Linnaeus 1753 (ICBN) B69
    |    `--*A. macrophthalmus Haswell 1880 B69
    |--Ambasia Boeck 1871 B69
    |    `--*A. atlantica (Milne Edwards 1830) B69 (see below for synonymy)
    |--Ambasiella Schellenberg 1935 B69
    |    `--*A. murmanica (Brüggen 1905) [=Ambasia murmanica] B69
    |--Eucallisoma Barnard 1961 B69
    |    `--*E. glandulosa Barnard 1961 B69
    |--Aristiopsis Barnard 1961 B69
    |    `--*A. tumidus Barnard 1961 B69
    |--Danaella Stephensen 1925 B69
    |    `--*D. mimonectes Stephensen 1925 B69
    |--Lepidepecreoides Barnard 1931 B69
    |    `--*L. xenopus Barnard 1931 B69
    |--Thoriella Stephensen 1915 B69
    |    `--*T. islandica Stephensen 1915 B69
    |--Crybelocyphocaris Shoemaker 1945 B69
    |    `--*C. tattersalli Shoemaker 1945 B69
    |--Centromedon Sars 1895 B69
    |    `--*C. pumilus (Liljeborg 1865) [=Anonyx pumilus] B69
    |--Azotostoma Barnard 1965 B69
    |    `--*A. fusta Barnard 1965 B69
    |--Bathyamaryllis Pirlot 1933 B69
    |    `--*B. perezii Pirlot 1933 B69
    |--Bathycallisoma Dahl 1959 B69
    |    `--*B. pacifica Dahl 1959 B69
    |--Chevreuxiella Stephensen 1915 B69
    |    `--*C. metopoides Stephensen 1915 B69
    |--Socarnoides Stebbing 1888 [incl. Acidostomella Schellenberg 1926] B69
    |    `--*S. kergueleni Stebbing 1888 B69
    |--Eurythenes Smith 1884 [incl. Euryporeia Sars 1895, Eurytenes Liljeborg 1865 (preoc.), Katius Chevreux 1905] B69
    |    `--*E. gryllus (Lichtenstein 1822) B69 [=Gammarus gryllus B69; incl. Lysianassa magellanica O01]
    |--Lysianella Sars 1882 B69
    |    `--*L. petalocera Sars 1882 B69
    |--Tryphosites Sars 1895 B69
    |    `--*T. longipes (Bate & Westwood 1863) B69 [=Anonyx longipes B69, Tryphosa longipes N00b]
    |--Alibrotus Milne Edwards 1840 (n. d.) B69
    |    `--*A. chauseicus (Milne Edwards 1830) [=Lysianassa chauseica] B69
    |--Cebocaris Barnard 1964 B69
    |    `--*C. grutesca Barnard 1964 B69
    |--Paralibrotus Stephensen 1923 B69
    |    `--*P. setosus Stephensen 1923 B69
    |--Pachynus Bulycheva 1955 B69
    |    `--*P. chelatum Bulycheva 1955 B69
    |--Koroga Holmes 1908 B69
    |    `--*K. megalops Holmes 1908 B69
    |--Gainella Chevreux 1911 B69
    |    `--*G. chelata Chevreux 1912 B69
    |--Paracallisomopsis Gurjanova 1962 B69
    |    `--*P. beljaevi Gurjanova 1962 B69
    |--Euonyx Norman 1867 [incl. Leptochela Boeck 1876 (preoc.)] B69
    |    `--*E. chelatus Norman 1867 B69 [incl. Opis leptochela Bate & Westwood 1869 N00b]
    |--Clepidecrella Barnard 1962 B69
    |    `--*C. cabinda Barnard 1962 B69
    |--Crybelocephalus Tattersall 1906 B69
    |    `--*C. megalurus Tattersall 1906 B69
    |--Cyclocaris Stebbing 1888 B69
    |    |--*C. tahitensis Stebbing 1888 B69
    |    `--C. faroensis Norman 1900 N00b
    |--Derjugiana Gurjanova 1962 B69
    |    `--*D. insolita Gurjanova 1962 B69
    |--Neoambasia Dahl 1959 B69
    |    `--*N. tumicornis (Nicholls 1938) [=Ambasiopsis tumicornis] B69
    |--Paracyphocaris Chevreux 1905 B69
    |    `--*P. praedator Chevreux 1905 B69
    |--Glycerina Haswell 1882 [incl. Glycera Haswell 1880 non Savigny 1818] B69
    |    `--*G. tenuicornis (Haswell 1880) [=Glycera tenuicornis] B69
    |--Scopelocheiropsis Schellenberg 1926 B69
    |    `--*S. abyssalis Schellenberg 1926 B69
    |--Ensayara Barnard 1964 B69
    |    `--*E. ramonella Barnard 1964 B69
    |--Figorella Barnard 1962 B69
    |    `--*F. tanidea Barnard 1962 B69
    |--Metacyphocaris Tattersall 1906 B69
    |    `--*M. helgae Tattersall 1906 B69
    |--Kyska Shoemaker 1964 B69
    |    `--*K. dalli Shoemaker 1964 B69
    |--Lepidepecreopsis Stephensen 1925 B69
    |    `--*L. biloba Stephensen 1925 B69
    |--Menigratopsis Dahl 1945 B69
    |    `--*M. svenillsoni Dahl 1945 B69
    |--Mesocyclocaris Birstein & Vinogradov 1964 B69
    |    `--*M. gracilis Birstein & Vinogradov 1964 B69
    |--Mesocyphocaris Birstein & Vinogradov 1960 B69
    |    `--*M. longicaudatus Birstein & Vinogradov 1960 B69
    |--Metacyclocaris Birstein & Vinogradov 1955 B69
    |    `--*M. polycheles Birstein & Vinogradov 1955 B69
    |--Metambasia Stephensen 1923 B69
    |    `--*M. faeroensis Stephensen 1923 B69
    |--Microlysias Stebbing 1918 B69
    |    `--*M. xenokeras Stebbing 1918 B69
    |--Ocosingo Barnard 1964 B69
    |    `--*O. borlus Barnard 1964 B69
    |--Paracallisoma Chevreux 1903 B69
    |    `--*P. alberti Chevreux 1903 B69
    |--Paronesimoides Pirlot 1933 B69
    |    `--*P. lignivorus Pirlot 1933 B69
    |--Paronesimus Stebbing 1894 B69
    |    `--*P. barentsi Stebbing 1894 B69
    |--Phoxostoma Barnard 1925 B69
    |    `--*P. algoense Barnard 1925 B69
    |--Podoprionella Sars 1895 B69
    |    `--*P. norvegica Sars 1895 B69
    |--Podoprionides Walker 1906 B69
    |    `--*P. incerta Walker 1906 B69
    |--Prachynella Barnard 1964 B69
    |    `--*P. lodo Barnard 1964 B69
    |--Procyphocaris Barnard 1961 B69
    |    `--*P. primata Barnard 1961 B69
    |--Pseudambasia Stephensen 1927 B69
    |    `--*P. bipartita Stephensen 1927 B69
    |--Pseudoanonyx Kudrjaschov 1965 B69
    |    `--*P. caecus Kudrjaschov 1965 B69
    |--Rifcus Kudrjaschov 1965 B69
    |    `--*R. auspicatus Kudrjaschov 1965 B69
    |--Shackletonia Barnard 1931 B69
    |    `--*S. robusta Barnard 1931 B69
    |--Socarnella Walker 1904 B69
    |    `--*S. bonnieri Walker 1904 B69
    |--Stephensenia Schellenberg 1928 B69
    |    `--*S. haematopus Schellenberg 1928 B69
    |--Tryphosoides Schellenberg 1931 B69
    |    `--*T. falcata Schellenberg 1931 B69
    |--Vijaya Walker 1904 B69
    |    `--*V. tenuipes Walker 1904 B69
    |--Onisimus Boeck 1871 [=Pseudalibrotus Della Valle 1893] B69
    |    |--*O. litoralis (Krøyer 1845) B69 [=Anonyx litoralis B69, Alibrotus littoralis N00b, Pseudalibrotus litoralis B69]
    |    |--O. brevicaudatus S00
    |    `--O. plautus (Krøyer 1845) [=Anonyx plautus] B69
    |--Ichnopus Costa 1853 B69
    |    |--*I. taurus Costa 1853 B69
    |    `--I. spinicornis Boeck 1861 BD95 [incl. I. calceolatus Heller 1867 N00a]
    |--Lepidepecreum Bate & Westwood 1868 B69
    |    |--*L. longicorne (Bate & Westwood 1861) B69, BD95 (see below for synonymy)
    |    `--L. umbo B69
    |--Acidostoma Liljeborg 1865 B69
    |    |--*A. obesum (Bate & Westwood 1861) B69, BD95 [=Anonyx obesus B69]
    |    `--A. nodiferum Stephensen 1923 [incl. A. sarsi (Lincoln 1979)] BD95
    |--Ambasiopsis Barnard 1931 B69
    |    |--*A. georgiensis Barnard 1931 B69
    |    `--A. faeroensis N38
    |--Paralicella Chevreux 1908 B69
    |    |--*P. tenuipes Chevreux 1908 B69
    |    `--P. scotiae (Chilton 1912) [=Alicella scotiae] B69
    |--Menigrates Boeck 1871 B69
    |    |--*M. obtusifrons (Boeck 1861) B69 [=Anonyx obtusifrons B69; incl. A. brachycercus Lilljeborg 1865 N00b]
    |    `--M. angustipes Gurjanova 1962 B69
    |--Kerguelenia Stebbing 1888 B69
    |    |--*K. compacta Stebbing 1888 B69
    |    `--K. borealis Sars 1895 B69
    |--Onesimoides Stebbing 1888 B69
    |    |--*O. carinatus Stebbing 1888 B69
    |    `--O. chelatus B69
    |--Parawaldeckia Stebbing 1910 B69
    |    |--*P. thomsoni (Stebbing 1906) [=Nannonyx thomsoni] B69
    |    `--P. kidderi B69
    |--Boeckosimus Barnard 1969 B69
    |    |--*B. edwardsii (Krøyer 1846) [=Anonyx edwardsii, Onisimus edwardsii] B69
    |    `--B. normani (Sars 1895) [=Onisimus normani] B69
    |--Valettiopsis Holmes 1908 B69
    |    |--*V. dentatus Holmes 1908 B69
    |    `--V. multidentata Barnard 1961 B69
    |--Hirondellea Chevreux 1889 [incl. Tetronychia Stephensen 1923] B69
    |    |--*H. trioculata Chevreux 1889 B69
    |    `--H. brevicaudata Chevreux 1910 B69
    |--Pseudokoroga Schellenberg 1931 B69
    |    |--*P. barnardi Schellenberg 1931 B69
    |    `--P. rima B69
    |--Paracentromedon Chevreux & Fage 1925 B69
    |    |--*P. crenulatum (Chevreux 1900) [=Centromedon crenulatum] B69
    |    `--P. manene HS01
    |--Sophrosyne Stebbing 1888 [incl. Paropisa Stebbing 1899] B69
    |    |--*S. murrayi Stebbing 1888 B69
    |    `--S. robertsoni Stebbing & Robertson 1891 B69
    |--Socarnes Boeck 1871 B69
    |    |--*S. vahlii (Krøyer 1838) B69 [=Lysianassa vahlii B69, Anonyx vahlii N00a]
    |    |--S. erythrophthalmus Robertson 1892 N00a
    |    |--S. filicornis Heller 1866 BD95
    |    `--S. obesus (Chevreux 1927) [=Socarnopsis obesa] B69
    |--Pachychelium Stephensen 1925 B69
    |    |--*P. davidis Stephensen 1925 B69
    |    |--P. antarcticum N38
    |    `--P. oculatum N38
    |--Nannonyx Sars 1895 B69
    |    |--*N. goesii (Boeck 1871) B69 [=Orchomene goesii B69, Orchemene goesii N00b]
    |    |--N. propinquus Chevreux 1911 BD95
    |    `--N. spinimanus Walker 1895 BD95
    |--Parambasia Walker & Scott 1903 B69
    |    |--*P. forbesi Walker & Scott 1903 B69
    |    |--P. anomala (Nicholls 1938) [=Lysianassa anomala] B69
    |    `--P. rossi Stephensen 1927 B69
    |--Stomacontion Stebbing 1899 B69
    |    |--*S. pepinii (Stebbing 1888) [=Acontiostoma pepinii] B69
    |    |--S. capense Barnard 1916 B69
    |    `--S. insigne Barnard 1932 B69
    |--Normanion Bonnier 1893 [=Normania Boeck 1871 non Brady 1866] B69
    |    |--*N. quadrimana (Bate & Westwood 1868) [=Opis quadrimana, *Normania quadrimana] B69
    |    |--N. abyssi B69
    |    `--N. sarsi Stebbing 1906 B69
    |--Tmetonyx Stebbing 1906 [incl. Hoplonyx Sars 1895 (preoc.)] B69
    |    |--*T. cicada (Fabricius 1780) B69 (see below for synonymy)
    |    |--T. exiguus (Chevreux 1902) BD95
    |    |--‘Haplonyx’ leucophthalmus Sars 1891 N00b
    |    |--T. nardonis (Heller 1866) BD95
    |    |--T. nobilis B69
    |    `--‘Haplonyx’ similis Sars 1891 N00b
    |--Schisturella Norman 1900 B69
    |    |--*S. pulchra (Hansen 1887) [=Tryphosa pulchra] B69
    |    |--S. abyssi (Chevreux 1926) [=Pseudonesimus abyssi] B69
    |    |--S. adversicola (Barnard 1925) [=Lakota adversicola] B69
    |    `--S. rotundata (Barnard 1925) [=Lakota rotundatus] B69
    |--Cheirimedon Stebbing 1888 B69
    |    |--*C. crenatipalmatus Stebbing 1888 B69
    |    |--C. femoratus [incl. C. dentimanus Chevreux 1906] B69
    |    |--C. fougneri Walker 1903 B69
    |    |--C. latimanus Sars 1895 B69
    |    `--C. pectinipalma Barnard 1925 B69
    |--Anonyx Krøyer 1838 [incl. Chironesimus Sars 1895, Lakota Holmes 1908] B69
    |    |--*A. nugax (Phipps 1774) B69 (see below for synonymy)
    |    |--A. cicadoides Stebbing 1888 B69
    |    |--A. debruynii B69 [=Chironesimus debruynii S00]
    |    |--A. gularis CS77
    |    `--A. lilljeborgii S00
    |--Tryphosella Bonnier 1893 B69
    |    |--*T. sarsi Bonnier 1893 B69
    |    |--T. abyssi Norman 1900 N00b
    |    |--T. angulata N00b
    |    |--T. barbatipes (Stebbing 1888) [=Tryphosa barbatipes, Uristes barbatipes] B69
    |    |--T. coxalis (Barnard 1962) [=Tryphosites coxalis] B69
    |    |--T. georgiana [incl. T. albina] N38
    |    |--T. hoeringii (Boeck 1870) [=Tryphosa hoeringii, Tryphosa hoerringii, Tryphosella hoerringii] N00b
    |    |--T. minima (Chevreux 1911) BD95
    |    `--T. nanoides (Lilljeborg 1865) [=Anonyx nanoides, Tryphosa nanoides] N00b
    |--Lepidepecreella Schellenberg 1926 [incl. Paracyclocaris Barnard 1930] B69
    |    |--*L. ctenophora Schellenberg 1926 B69
    |    |--L. bidens N38
    |    |--L. cymba N38
    |    |--L. emarginata Nicholls 1938 N38
    |    `--L. ovalis N38
    |--Hippomedon Boeck 1871 [incl. Elimedon Barnard 1962, Paratryphosites Stebbing 1899, Platamon Stebbing 1888] B69
    |    |--*H. holbolli (Krøyer 1846) [=Anonyx holbolli] B69
    |    |--H. abyssi (Goës 1866) [=Lysianassa abyssi, Paratryphosites abyssi] N00b
    |    |--H. denticulatus (Bate 1857) BD95 [=Anonyx denticulatus N00b]
    |    |--H. kergueleni PG98
    |    |--‘Platamon’ longimanus N00b
    |    |--H. massiliensis Bellan-Santini 1965 BD95
    |    |--H. oculatus Chevreux & Fage 1925 BD95
    |    `--H. propinquus Sars 1890 N00b
    |--Lysianassa Milne Edwards 1830 (see below for synonymy) B69
    |    |--*L. costae Milne Edwards 1830 B69
    |    |--L. alba (Holmes 1903) [=*Lysianopsis alba] B69
    |    |--L. bispinosa Della Valle 1893 BD95
    |    |--L. ceratina (Walker 1889) BD95 [=Lysianax ceratinus N00a]
    |    |--‘Lysianax’ cubensis Stebbing 1897 B69
    |    |--L. heterodonta (Pirlot 1936) [=*Arugella heterodonta] B69
    |    |--L. hypocrita B69
    |    |--L. insperata Lincoln 1979 BD95
    |    |--L. longicornis Lucas 1846 E12 [=Lysianax longicornis N00a]
    |    |--‘*Pronannonyx’ minimus Schellenberg 1953 B69
    |    |--L. nasuta Dana 1853 [=*Shoemakerella nasuta] B69
    |    |--‘*Aruga’ oculata Holmes 1908 B69
    |    |--L. pilicornis Heller 1866 BD95
    |    |--L. plumosa Boeck 1871 BD95
    |    `--‘Lysianax’ septentrionalis Della Valle 1893 N00a
    |--Podoprion Chevreux 1891 BD95, B69 [Podoprionidae]
    |    `--*P. bolivari Chevreux 1891 B69
    |--Scopelocheirus Bate 1857 BD95, B69 [incl. Callisoma Costa 1851 non Agassiz 1846 B69; Scopelocheiridae]
    |    |--*S. crenatus Bate 1857 B69 [=Callisoma crenata N00b]
    |    |--S. hopei (Costa 1851) BD95 [=*Callisoma hopei B69]
    |    `--‘Anonyx’ kroeyeri Bruzelius 1859 [=Callisoma kroeyeri] N00b
    |--Aristias Boeck 1871 BD95, B69 [Aristiidae]
    |    |--*A. tumidus (Krøyer 1846) [=Anonyx tumidus] B69
    |    |--A. antarcticus N38
    |    |--A. collinus N38
    |    `--A. neglectus Hansen 1887 N00b
    `--Orchomene Boeck 1871 [incl. Orchomenella Sars 1891, Orchomenopsis Sars 1895, Tryphosa Boeck 1871] B69
         |--*O. serrata (Boeck 1861) [=Anonyx serratus] B69
         |--‘Orchomenella’ abyssorum [=O. chilensis f. abyssorum] N38
         |--‘Tryphosa’ adarei N38
         |--O. amblyops Sars 1895 B69
         |--O. batei Sars 1895 B69
         |--‘Tryphosa’ carinata Schellenberg 1926 B69
         |--O. cavimanus [=Orchomenella cavimanus; incl. Orchomenopsis zschauii] N38
         |--‘Orchomenella’ franklini N38
         |--‘Orchomenella’ groenlandica B69
         |--O. humilis (Costa 1853) [=Lysianassa humilis, Anonyx humilis; incl. A. melanophthalmus Norman 1867] N00b
         |--‘Tryphosa’ kergueleni N38
         |--‘Orchomenella’ macronyx N38
         |--O. minuta (Krøyer 1846) B69 [=Anonyx minutus B69, *Orchomenella minuta B69, Tryphosa minuta N00b]
         |--‘Tryphosa’ murrayi N38
         |--‘Anonyx’ nanus Krøyer 1846 B69 (see below for synonymy)
         |--‘Orchomenella’ nodimanus [=Orchomenopsis nodimanus] N38
         |--O. obtusa (Sars 1895) B69
         |--O. pectinatus Sars 1882 N00b
         |--O. pinguis (Boeck 1860) [=Anonyx pinguis, Orchemenella pinguis, Tryphosa pinguis] N00b
         |--‘Tryphosa’ pusilla Sars 1885 N00b
         |--‘Orchomenella’ rossi [=Orchomenopsis rossi, Orchomenella chilensis f. rossi] N38
         |--O. takoradia Barnard 1961 B69
         `--‘Tryphosa’ triangularis N38

*Ambasia atlantica (Milne Edwards 1830) B69 [=Gammarus atlanticus B69, Lysianassa atlantica N00a; incl. Ambasia danielssenii Boeck 1870 N00a]

‘Anonyx’ nanus Krøyer 1846 B69 [=Orchemenella nana N00b, *Tryphosa nana B69; incl. T. ciliata Sars 1882 N00b, Orchomenella ciliata B69]

*Anonyx nugax (Phipps 1774) B69 [=Cancer nugax B69; incl. A. appendiculata Kröyer 1838 N00b, A. lagena Kröyer 1838 N00b]

*Lepidepecreum longicorne (Bate & Westwood 1861) B69, BD95 [=Anonyx longicornis B69; incl. L. carinatum Bate & Westwood 1869 N00b, L. mirabile Meinert 1890 N00b]

Lysianassa Milne Edwards 1830 [incl. Aruga Holmes 1908, Arugella Pirlot 1936, Lysianassina Costa 1867, Lysianax Stebbing 1888, Lysianopsis Holmes 1903, Pronannonyx Schellenberg 1953, Shoemakerella Pirlot 1936] B69

*Perrierella audouiniana (Bate 1857) B69 [=Lysianassa audouiniana B69, Aristias audouinianus N00b, Pararistias auduoinianus N00b; incl. Perrierella crassipes Chevreux & Bouvier 1892 N00b]

*Tmetonyx cicada (Fabricius 1780) B69 [=Oniscus cicada B69, Haplonyx cicada N00b; incl. Anonyx gulosus Kröyer 1844 N00b, A. norvegicus Lilljeborg 1851 N00b]

*Type species of generic name indicated

REFERENCES

[BD95] Bakalem, A., & J.-C. Dauvin. 1995. Inventaire des crustacés amphipodes (Gammaridea, Caprellidea, Hyperiidea) des costes d'Algérie: essai de synthèse. Mésogée 54: 49–62.

[B69] Barnard, J. L. 1969. The families and genera of marine gammaridean Amphipoda. United States National Museum Bulletin 271: i–vi, 1–535.

[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[N38] Nicholls, G. E. 1938. Australasian Antarctic Expedition 1911–1914. Under the Leadership of Sir Douglas Mawson, O.B.E., B.E., D.Sc., F. R. S. Scientific Reports. Series C—Zoology and Botany vol. 2 pt 4. Amphipoda Gammaridea. David Harold Paisley, Government Printer: Sydney (Australia).

[N00a] Norman, A. M. 1900a. British Amphipoda of the tribe Hyperiidea and the families Orchestiidae and some Lysianassidae. Annals and Magazine of Natural History, series 7, 5: 126–144.

[N00b] Norman, A. M. 1900b. British Amphipoda: fam. Lysianassidae (concluded). Annals and Magazine of Natural History, series 7, 5: 196–214, pl. 6.

[O01] Ohlin, A. 1901. On a new "bipolar" schizopod. Annals and Magazine of Natural History, series 7, 7: 371–374.

[PG98] Probert, P. K., & S. L. Grove. 1998. Macrobenthic assemblages of the continental shelf and upper slope off the west coast of South Island, New Zealand. Journal of the Royal Society of New Zealand 28: 259–280.

[S00] Stebbing, T. R. R. 1900. Arctic Crustacea: Bruce collection. Annals and Magazine of Natural History, series 7, 5: 1–16.

[S86] Stock, J. H. 1986. Order Amphipoda: an introduction. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 494–496. E. J. Brill/Dr W. Backhuys: Leiden.