Belongs within: Phytophaga.
Contains: Donacia, Sagrinae, Colaspoides, Colasposoma, Corynodes, Nodostoma, Colaspis, Hispinae, Crioceris, Lema, Acanthoscelides, Bruchus, Cassida, Cryptocephalini, Clytrini, Galerucinae, Chrysomelini.
The Chrysomelidae, leaf beetles, are a cosmopolitan family of beetles that mostly feed on living plant tissue. Adults feed externally whereas larvae may feed either externally or internally. Subgroups of the Chrysomelidae include the Bruchinae, seed-feeding species with a somewhat rostrate head that superficially resemble weevils in appearance. The Chrysomelinae are generally ovate beetles characterised by a truncate clypeus whose larvae feed externally and are often armed with repugnatorial glands. The Cryptocephalinae usually have a strongly hypognathous head that is deeply inserted into the prothorax. The Eumolpinae have larvae that are found in soil and probably feed on plant roots (Lawrence & Britton 1991).
Chrysomelidae
|--+--Criocerinae [Crioceridae] MT11
| | |--Crioceris MW15
| | |--Stethopachys formosa LB91
| | |--Lema LB91
| | `--Lilioceris B14
| | |--L. cheni Gressit & Kimoto 1961 B14
| | |--L. egena B14
| | |--L. lilii B14
| | `--L. nigripes LB91
| `--Bruchinae [Bruchidae, Lariidae, Mylabridae] MW15
| |--Caryedes brasiliensis GE05
| |--Caryoborus pallidus B88
| |--Acanthoscelides SJ73
| |--Spermophagus cardui C01
| |--Bruchus MW15
| |--Sennius morosus G05
| |--Mesolaria longala Zhang 1986 RJ93
| |--Caryedon B14
| | |--C. abdominalis B14
| | `--C. serratus (Olivier 1790) B14
| |--Rhaebus [Rhaebini] B14
| | |--R. ammoni B14
| | `--R. mannerheimi Motschulsky 1845 B14
| |--Callosobruchus LB91
| | |--C. chinensis N57
| | `--C. maculatus RCT07
| |--Stator G05
| | |--S. limbatus G05
| | `--S. pruininus G05
| |--Pachymerus RD77
| | |--P. chinensis RD77
| | `--P. dactris O49
| `--Bruchidius F05
| |--B. incarnatus TG96
| |--B. lyndhurstensis LB91
| `--B. niger Anton 2004 F05
`--+--+--Cassidinae [Cassidini] MW15
| | |--Cassida B88
| | |--Hemisphaerota cyanea (Say 1824) B14
| | |--Meroscalcis selecta LB91
| | |--Aspidomorpha LB91
| | |--Brontispa B14
| | | |--B. longissima (Gestro 1885) B14
| | | `--B. simonthomasi Gressitt 1960 B14
| | |--Dicladispa B14
| | | |--D. armigera (Olivier 1808) B14
| | | `--D. birendra Maulik 1919 B14
| | |--Acromis B14
| | | |--A. sparsa (Boheman 1854) B14
| | | |--A. spinifex B14
| | | `--A. venosa B14
| | `--Cephaloleia belti B14
| `--Cryptocephalinae [Camptosomata, Clytrinae] MW15
| | i. s.: Aporocera ZS10
| | Fulcidax monstrosa (Fabricius 1798) B14
| | Leasia LB91
| | Neochlamisus bebbiana MW15
| | Lachnabothra QI93
| | Prasonotus QI93
| | Loxopleurus B70
| |--Cryptocephalini SW07
| |--Clytrini SW07
| `--Chlamisus [Chlamisini] LB91
`--+--Galerucinae MW15
`--Chrysomelinae MW15
| i. s.: Leptinotarsa B14
| |--L. decemlineata Say 1824 B14
| `--L. juncta B14
| Sterromela trimaculata LB91
| Phratora B14
| | i. s.: P. frosti B14
| | P. hudsonica B14
| `--P. (Phyllodecta) B14
| |--‘Phyllodecta’ longula C01
| |--P. (P.) polaris (Schneider 1886) B14
| |--‘Phyllodecta’ vitellinae C01
| `--‘Phyllodecta’ vulgatissima [incl. P. vulgatissima var. obscura] C01
| Chlamys L02
| |--C. cristata [=Cryptocephalus (Chlamys) cristatus] G20
| |--C. formosana Bates 1866 B66
| |--C. gibba [=Clythra gibba] L02
| |--C. monstrosa [=Clythra monstrosa] L02
| |--C. plicata [=Clythra plicata] L02
| `--C. tuberosa L02
| Orina RD77
| Phaedon RD77
| |--P. armoraciae C01
| `--P. cochleariae H91
| Phytodecta RD77
| |--P. chrysomeloides Jacoby 1889 J89a
| |--P. formicata R13
| |--P. linnaeana (see below for synonymy) C01
| |--P. rufipes C01
| `--P. viminalis R13
| Zygogramma piceicollis Stål 1859 Lu89
|--Chrysomelini MW15
`--Timarcha [Timarchini] B14
|--T. punica Lucas 1847 E12
`--T. tenebricosa (Fabricius 1775) B14
Chrysomelidae incertae sedis:
Pseudomegamerus grandis GE05, P02
Eucolaspis brunnea WFS04
Oreina cacaliae AM07
Arnomus brouni T27
Nesaecrepidia BHM08
Protoscelis GE05
Protosceloides GE05
Eurydemus oculatus Chap. 1879 G89
Physimera DL99
Cheilotoma musciformis F05
|--C. m. musciformis F05
`--C. m. hispanica Medvedev 2004 F05
Haemonia WP99
Gastrophysa viridula V09
Pachybrachys C01
|--P. fimbriolatus C01
`--P. hieroghyphicus [incl. P. hieroghyphicus var. tristis] C01
Stylosomus flavus C01
Adoxus C01
|--A. obscurus C01
`--A. villosulus C01
Gastroidea C01
|--G. polygoni [incl. G. polygoni var. ruficollis] C01
`--G. viridula C01
Entomoscelis C01
|--E. adonidis C01
`--E. orinalis F92
Xenomela heydeni C01
Paropsides C01
|--P. duodecimpustulatus C01
`--P. nigrofasciatus Jac. 1888 F89
Hydrothassa glabra C01
Sternoplatys weisei Csiki 1901 C01
Melasoma C01
|--M. collaris [incl. M. collaris var. daurica] C01
|--M. lapponica C01
|--M. populi C01
`--M. tremulae R13
Malacosoma Chv. 1846 [incl. Exocera] FS90
|--M. collaris C01
`--M. detritum Fab. 1801 [incl. M. guadeloupense, M. pallidicorne] FS90
Phyllobrotica C01
|--P. quadrimaculata B56
|--P. signata C01
`--P. unicolor J89a
Lochmaea C01
|--L. birmanica Jacoby 1889 J89a
|--L. capreae C01
|--L. rugosa [=Galerucella rugosa] J89a
`--L. suturalis F92
Adimonia C01
|--A. incisicollis C01
|--A. mongolica Csiki 1901 C01
|--A. pomonae C01
|--A. sedakovi C01
|--A. tanaceti C01
`--A. violacea Lucas 1849 E12
Pallasia absinthii C01
Ochrosis salicariae C01
Chalcoides C01
|--C. aurata C01
|--C. cyanea C01
|--C. helxines [incl. C. helxines var. fulvicornis] C01
`--C. metallica C01
Mantura rustica [incl. M. rustica var. suturalis] C01
Capraita B14
Platymela sticticollis B14
Spilopyra [Spilopyrinae] B14
`--S. sumptuosa Baly 1860 B14
Syneta [Synetinae] B14
|--S. adamsi B14
`--S. betulae (Fabricius 1792) B14
Zeugophora [Zeugophorinae] B14
| i. s.: Z. flavicollis C01
`--Z. (Zeugophora) B14
|--Z. (Z.) scutellaris Suffrian 1840 B14
`--Z. (Z.) subspinosa B14
Lamprosomatinae [Lamprosomatini] B14
|--Oomorphus B14
| |--O. concolor (Sturm 1807) B14
| `--O. japanus B14
`--Lamprosoma B66
|--L. alienum Bates 1866 B66
|--L. concolor B66
`--L. cupricolle L90a
Donacia B14
Sagrinae B14
Eumolpinae [Eumolpidae, Pseudocolaspides] MT11
|--Eka M30
|--Trichochrysea hirta (Fabricius 1801) MT11
|--Malegia Lefevre 1883 MT11
|--Colaspoides MT11
|--Cleorina aeneomicans (Baly 1867) [=Nodostoma aeneomicans; incl. C. aulica Lef. 1885] Le89a
|--Colasposoma MT11
|--Phytorus dilatatus Jacoby 1884 MT11
|--Apolepis MT11
|--Lepidocolaspis MT11
|--Chloropterus B14
|--Edusella abdominalis LB91
|--Geloptera LB91
|--Tomyris LB91
|--Cleoporus badius Lefèvre 1889 Le89a
|--Corynodes Le89a
|--Nodostoma Le89a
|--Dicolectes aulicus Lef. 1886 Le89b
|--Eryxia baikiei Baly 1865 Le89b
|--Colaspis L90a
|--Sphaeropis humeralis Lef. 1878 L90a
|--Phanaeta ruficollis Lef. 1878 L90a
|--Iphimeis L90b
|--Chalcophana cyanipennis Lefèvre 1890 L90b
|--Agbalus violaceus [=Corysthea violacea] L90b
|--‘Nephus’ Jacoby 1889 non Mulsant 1846 J89b
| `--*N. femoratus Jacoby 1889 J89b
|--Chalcolema Jacoby 1890 MT11
| `--C. fulvicornis (Jacoby 1889) [=Platycorynus fulvicornis] MT11
|--Osnaparis Fairmaire 1889 F89
| `--*O. nucea Fairmaire 1889 F89
|--Leprotoides MT11
| |--L. flavipes Jacoby 1896 MT11
| `--L. hitamtitik Medvedev & Takizawa 2011 MT11
|--Tricliona MT11
| |--T. oculata Medvedev & Takizawa 2011 MT11
| `--T. sulcatipennis Jacoby 1896 MT11
|--Pagria MT11
| |--P. flavopustulata Baly 1874 MT11
| `--P. sumatrensis Lefevre 1887 MT11
|--Aulacolepis Le89a
| |--A. decorata J89a
| `--A. mouhoti Baly 1863 Le89a
|--Mouhotina Le89a
| |--M. rufipes Lefèvre 1889 Le89a
| `--M. tibialis (Lef. 1885) [=Stethotes tibialis] Le89a
|--Nerissus Le89b
| |--N. affinis Lefèvre 1889 Le89b
| `--N. strigosus Chap. 1874 Le89b
|--Syagrus Le89b
| |--S. alluaudi Lefèvre 1889 Le89b
| `--S. calcaratus (Fabr. 1775) (see below for synonymy) Le89b
|--Menius Le89b
| |--M. aeratus [=Syagrus aeratus] Le89b
| `--M. tarsalis Lef. 1887 Le89b
|--Lamprosphaerus L90a
| |--L. caeruleatus Lef. 1877 L90a
| `--L. minutus [=Chrysodina minuta] L90a
|--Alethaxius L90b
| |--A. brevis Lefèvre 1890 L90b
| |--A. parvulus Lefèvre 1889 Le89c
| `--A. tuberculifer Lefèvre 1890 L90b
|--Corysthea L90b
| |--C. ruficollis Lefèvre 1890 L90b
| `--C. rugulosa Lefèvre 1890 L90b
|--Endocephalus L90b
| |--E. fasciatus Lefèvre 1890 L90b
| `--E. lefevrei Harold 1874 L90b
|--Xanthonia Baly 1863 [incl. Icogramma Weise 1922] MT11
| |--X. insularis Medvedev & Takizawa 2011 MT11
| |--X. lineigera (Weise 1922) MT11
| `--X. minuta Pic 1929 MT11
|--Aulexis MT11
| |--A. elongata J89a
| |--A. pallida (Lefevre 1887) MT11
| `--A. tibialis Jacoby 1889 J89a
|--Basilepta MT11
| |--B. flavescens (Motschulsky 1866) [incl. Nodostoma beccarii Jacoby 1884] MT11
| |--B. nigromarginata (Jacoby 1896) MT11
| `--B. simplex (Jacoby 1884) MT11
|--Pseudocolaspis Le89b
| |--P. albostriata G89
| |--P. setosa Lucas 1846 E12
| `--P. setulosa Lef. 1887 Le89b
|--Choris Lefèvre 1877 [incl. Mevania Harold 1874 non Stäl 1865] L90a
| |--C. kirschi (Har. 1874) [=Mevania kirschi] L90a
| |--C. lateralis Lef. 1877 L90a
| `--C. nucea Lef. 1877 L90a
|--Rhabdopterus L90a
| |--R. circumdatus Lefèvre 1889 Le89c
| |--R. cuprinus Lefèvre 1890 L90a
| |--R. fulvipes L90a
| `--R. obsitus Lef. 1885 L90a
|--Typophorus L90a
| |--T. annulatus Lef. 1877 L90a
| |--T. exilis Lef. 1878 L90a
| `--T. viridicyaneus (Crotch 1873) [=Paria viridicyanea; incl. T. sturmi Lef. 1877] L90a
|--Abirus MT11
| |--A. flavopilosus Jacoby 1884 MT11
| |--A. fortunei (Baly 1861) [=Dermorhytis fortunei] F89
| |--A. harmandi Lef. 1876 Le89a
| `--A. piceipes J89b
|--Scelodonta MT11
| |--S. dillwyni (Stephens 1831) MT11
| |--S. nitidula Baly 1864 Le89a
| |--S. strigicollis J89a
| `--S. vittata [=Eumolpus vittatus, Heteraspis vittata] Le89a
|--Chrysochus B14
| |--C. asclepiadeus C01
| |--C. auratus (Fabricius 1775) B14
| |--C. cobaltinus B14
| `--C. pulcher J89a
|--Chrysodina L90a
| |--C. cribricollis Lefèvre 1890 L90a
| |--C. laevicollis Lefèvre 1885 L90a
| |--C. parvula Lef. 1885 L90a
| `--C. tibialis Lefèvre 1890 L90a
|--Nodonota L90a
| |--N. laeta Lef. 1878 L90a
| |--N. simoni Lefèvre 1890 L90a
| |--N. singularis Lefèvre 1890 L90a
| `--N. tuberculata Lef. 1875 L90a
|--Ephyraea Lefèvre 1890 L90a
| |--E. castanea Lefèvre 1890 L90a
| |--E. chapuisi (Har. 1874) [=Chrysodina chapuisi] L90a
| |--E. glaucescens Lefèvre 1890 L90a
| `--E. hypomelas Lefèvre 1890 L90a
|--Talurus Lefèvre 1890 L90b
| |--*T. rugosus [=Rhabdopterus rugosus] L90b
| |--T. fulgens Lefèvre 1890 L90b
| |--T. picitarsis Lefèvre 1890 L90b
| `--T. salvini [=Rhabdopterus salvini] L90b
|--Aoria Baly 1863 MT11
| |--A. bowringii (Baly 1860) [=Adoxus bowringii] Le89a
| |--A. nigripes (Baly 1860) [=Adoxus nigripes] Le89a
| |--A. pusilla Lefèvre 1889 Le89a
| |--A. quinquemaculata Bates 1866 B66
| `--A. rufotestacea Fairmaire 1889 F89
|--Nodina MT11
| |--N. baliana Medvedev & Takizawa 2011 MT11
| |--N. chalcosoma J89a
| |--N. fulvitarsis Jacoby 1896 MT11
| |--N. pusilla Motsch. 1858 Le89a
| `--N. rotundata Motsch. 1858 Le89a
|--Rhyparida MT11
| |--R. armata Jacoby 1889 J89b
| |--R. bimaculata Jacoby 1889 J89a
| |--R. microdentata Medvedev & Takizawa 2011 MT11
| |--R. morosa B70
| `--R. nigrosignata Jacoby 1884 MT11
|--Callisina MT11
| |--C. fasciata Baly 1860 MT11
| | |--C. f. fasciata MT11
| | |--C. f. baliana Medvedev & Takizawa 2011 MT11
| | `--C. f. javana Medvedev & Takizawa 2011 MT11
| `--C. quadripustulata J89a
|--Eumolpus L02
| |--E. cyanellus Dejean in Boisduval 1835 B35
| |--E. pictus Perty 1832 [incl. Balya quadrimaculata Jacoby 1882] L90b
| |--E. pretiosus L02
| |--E. quattuorpunctatus L02
| |--E. surinamensis Fabr. 1775 L90a
| `--E. vitis [=Cryptocephalus (Eumolpus) vitis] G20
|--Pachnephorus MT11
| |--P. canus B14
| |--P. cylindricus Lucas 1846 E12
| |--P. porosus Baly 1878 MT11
| |--P. seriatus Lef. 1887 Le89a
| |--P. tessellatus (Duftschmid 1825) B14
| `--P. variegatus Lef. 1887 Le89a
|--Platycorynus Chevrolat 1837 MT11
| |--P. approximans Baly 1864 MT11
| |--P. fraternus (Baly 1964) MT11
| |--P. hijau Medvedev & Takizawa 2011 MT11
| |--P. lefevrei (Jacoby 1895) MT11
| |--P. rufescens Medvedev & Takizawa 2011 MT11
| `--P. rufipennis Pic 1934 MT11
|--Myochrous Le89c
| |--M. armatus Le89c
| `--M. curculionoides Lefèvre 1889 Le89c
`--Euryope hoehneli Lefèvre 1890 L90c
‘Aletes’ Chapuis 1874 nec Rafinesque 1815 nec Carpenter 1857 N47
Systena lherminieri [incl. S. pallida] FS90
Oxygona Chv. 1847 FS90
`--O. pallens Fab. 1792 [incl. O. modesta] FS90
Pseudodisonycha turquinensis Blake 1956 B56
Griburius larvatus AT01
Bassareus croceipennis AT01
Metachroma gracile Black 1937 B37
Megistops B37
|--M. adulta B37
|--M. fictor B37
|--M. lugubrina B37
|--M. ornatus B37
|--M. pretiosus B37
|--M. quattuornotata B37
|--M. rubropustulata B37
`--M. tabebuiae Blake 1937 B37
Paratrikona Spaeth 1923 B37
|--P. lerouxi [=Trikona lerouxi] B37
|--P. ovata Blake 1938 B38
|--P. turrifera [=Trikona turrifera] B37
`--P. turritella Blake 1937 B37
Batonota Hope 1839 B37
Akantaka Maulik 1916 B37
Trikona Maulik 1916 B37
`--T. humeralis B37
Elytrogona nigrodorsata Blake 1937 B37
Chalcosicya B38
|--C. costata Blake 1938 B38
|--C. ditrichota Blake 1938 B38
|--C. maestrensis B38
|--C. nana B38
`--C. rotunda Blake 1938 B38
Anisodera P92
Chalepus praeustus [=Odontota praeusta] FS90
Criocerina P92
Electrolema P92
Hadroscelus P92
Inclusus P92
Oposispa P92
Protanisodera P92
Sucinagonia P92
Profidia Gresset 1963 P92
`--*P. nitida Gresset 1963 P92
Palaeothona Jac. 1885 FS90
`--P. semifulva Jac. 1885 [incl. P. smaragdipennis] FS90
Euplectroscelis Crotch 1873 FS90
`--E. variabilis Jac. 1885 [incl. E. variabilis var. rubra Fleutiaux & Sallé 1890] FS90
Homophyla Har. 1877 FS90
`--H. krugi Weise 1885 FS90
Homophoeta Er. 1847 FS90
`--H. aequinoctialis Linné 1758 [incl. H. aequinoctialis var. albicollis Fab. 1787, H. leucospila] FS90
Cerotoma Chv. 1843 FS90
|--C. ruficornis Ol. 1791 [incl. C. decorata, C. denticornis] FS90
`--C. trifurcata WR-SG01
Melanispa Baly 1858 FS90
`--M. truncata Baly 1858 FS90
Mesomphalia Hope 1838 FS90
|--M. aenea Ol. 1790 FS90
`--M. exclamationis Linné 1767 FS90
Chelymorpha Boh. 1854 FS90
|--C. argus G17
|--C. cribraria Fab. 1775 [incl. C. apiata, C. multipunctata] FS90
`--C. polysticha Boh. 1854 FS90
Selenis Hope 1839 FS90
`--S. spinifex Linné 1763 [incl. S. cerastes] FS90
Omoplata Hope 1840 FS90
`--O. distincta Boh. 1854 FS90
Coptocycla Boh. 1855 FS90
|--C. bistripunctata Ol. 1790 [incl. C. aurichalcea, C. sexpunctata] FS90
|--C. cruciata Linné 1767 [incl. C. crux, C. cruciata var. fuliginosa, C. vicina] FS90
`--C. guadeloupensis Boh. 1855 [incl. C. gemmea] FS90
Anomalonyx Weise 1903 BP02
Hispinae MT11
Phytodecta linnaeana [incl. P. linnaeana var. decostigma, P. linnaeana var. nigricollis, P. linnaeana var. orientalis] C01
Syagrus calcaratus (Fabr. 1775) [=Cryptocephalus calcaratus, Eumolpus calcaratus; incl. Typophorus buqueti Dej. 1837, Syagrus buqueti, Brevicolaspis ruficollis Thomson 1858] Le89b
*Type species of generic name indicated
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