Showing posts with label Arecaceae. Show all posts
Showing posts with label Arecaceae. Show all posts

Coryphoideae

Chinese fan palm Livistona chinensis, copyright Forest and Kim Starr.


Belongs within: Arecaceae.
Contains: Licuala.

The Coryphoideae are a group of palms whose members often have palmate or costapalmate, induplicate leaves and/or apocarpous flowers.

<==Coryphoideae
    |--Borasseae HB09
    |    |--Hyphaeninae HB09
    |    `--Lataniinae HB09
    |         |--Latania doddigesii BP60
    |         `--Borassus flabellifer HB09, KJ05
    `--Corypheae BBB03
         |  i. s.: Guihaia BBB03
         |         Kerriodoxa BBB03
         `--Livistoninae BBB03
              |--Licuala SDK-L03
              |--Johannesteijsmannia altifrons SDK-L03, P88
              `--Livistona B12
                   |--L. alfredii MLP09
                   |--L. australis (Brown) von Martius 1838 HE80 [=Corypha australis B78]
                   |--L. benthamii B00
                   |--L. chinensis A80b
                   |    |--L. c. var. chinensis A80b
                   |    |--L. c. var. boninensis A80a
                   |    `--L. c. var. subglobosa A80b
                   |--L. decipiens B00
                   |--L. drudei B00
                   |--L. eastonii LK14
                   |--L. humilis [incl. L. leichhardtii] B78
                   |--L. inermis B78
                   |--L. lorophylla B12
                   |--L. mariae B00
                   |--L. muelleri B00
                   |--L. nasmophila B12
                   `--L. nitida B00

Coryphoideae incertae sedis:
  Chamaerops humilis Z03, R-CT01
  Copernicia Z03
    |--C. alba F11
    `--C. australis F11
  Chuniophoenix Z03
  Phoenix Z03
    |--P. canariensis Chabaud 1882 HE80
    |--P. dactylifera MLP09
    |--P. reclinata DS04
    |--P. sylvestris KJ05
    `--P. theophrasti D81
  Medemia Z03
  Satranalia Z03
  Thrinax F11 [Thrinacinae SDK-L03]
    `--T. radiata F11

*Type species of generic name indicated

REFERENCES

[A80a] Aoki, J. 1980a. A revision of the oribatid mites of Japan. I. The families Phthiracaridae and Oribotritiidae. Bulletin of the Institute of Environmental Science and Technology, Yokohama National University 6 (2): 1–89.

[A80b] Aoki, J. 1980b. A revision of the oribatid mites of Japan. III. Families Protoplophoridae, Archoplophoridae and Mesoplophoridae. Proceedings of the Japanese Society of Systematic Zoology 18: 5–16.

[BP60] Baker, E. W., & A. E. Pritchard. 1960. The tetranychoid mites of Africa. Hilgardia 29 (11): 455–574.

[BBB03] Barfod, A. S., T. Burholt & F. Borchsenius. 2003. Contrasting pollination modes in three species of Licuala (Arecaceae: Coryphoideae). Telopea 10 (1): 207–223.

[B78] Bentham, G. 1878. Flora Australiensis: A description of the plants of the Australian Territory vol. 7. Roxburghiaceae to Filices. L. Reeve & Co.: London.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[B12] Braby, M. F. 2012. The butterflies of El Questro Wilderness Park, with remarks on the taxonomy of the Kimberley fauna, Australia. Records of the Western Australian Museum 27 (2): 161–175.

[DS04] Davis, J. I., D. W. Stevenson, G. Petersen, O. Seberg, L. M. Campbell, J. V. Freudenstein, D. H. Goldman, C. R. Hardy, F. A. Michelangeli, M. P. Simmons, C. D. Specht, F. Vergara-Silva & M. Gandolfo. 2004. A phylogeny of the monocots, as inferred from rbcL and atpA sequence variation, and a comparison of methods for calculating jackknife and bootstrap values. Systematic Botany 29 (3): 467–510.

[D81] Day, D. 1981. The Doomsday Book of Animals: A unique natural history of three hundred vanished species. Ebury Press: London.

[F11] Fraga, R. M. 2011. Family Icteridae (New World blackbirds). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 684–807. Lynx Edicions: Barcelona.

[HE80] Healy, A. J., & E. Edgar. 1980. Flora of New Zealand vol. 3. Adventive cyperaceous, petalous and spathaceous monocotyledons. P. D. Hasselberg, Government Printer: Wellington (New Zealand).

[HB09] Heatubun, C. D., W. J. Baker, J. P. Mogea, M. M. Harley, S. S. Tjitrosoedirdjo & J. Dransfield. 2009. A monograph of Cyrtostachys (Arecaceae). Kew Bulletin 64 (1): 67–94.

[KJ05] Katewa, S. S., A. Jain, B. L. Chaudhary & P. Galav. 2005. Some unreported medicinal uses of plants from the tribal area of Southern Rajasthan. Bulletin of the Botanical Survey of India 47: 121–130.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[MLP09] McKenzie, N. L., S. van Leeuwen & A. M. Pinder. 2009. Introduction to the Pilbara Biodiversity Survey, 2002–2007. Records of the Western Australian Museum Supplement 78 (1): 3–89.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[R-CT01] Ragusa-di Chiara, S., & H. Tsolakis. 2001. Phytoseiid faunas of natural and agricultural ecosystems in Sicily. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 522–529. CSIRO Publishing: Melbourne.

[SDK-L05] Saw, L. G., J. Dransfield & D. M. Keith-Lucas. 2003. Morphological diversity of the genus Licuala (Palmae). Telopea 10 (1): 187–206.

[Z03] Zona, S. 2003. Endosperm condition and the paradox of Ptychococcus paradoxus. Telopea 10 (1): 179–185.

Arecaceae

Nipah palm Nypa fruticans, copyright Tony Rodd.


Belongs within: Commelinidae.
Contains: Arecoideae, Coryphoideae, Calamoideae.

The Arecaceae, palms, are a group of mostly tropical and subtropical woody plants with large divided leaves. Most palms grow as an erect (usually unbranched) tree, but exceptions exist. In the nipa palm Nypa fruticans, found in mangrove habitats of the tropical Indo-Pacific region, the stem (which may be branching) grows lying on the substrate and only the leaves and flower stalks extend upwards. Fruits similar to those of Nypa are known from the fossil record going back to the Lower Paleocene. Foliage of palms is also known from the Cretaceous with Eolirion primigenium from the Lower Cretaceous of Austria representing the earliest record of the family (Collinson et al. 2003). Other basal genera of Arecaceae include Trithrinax, the caranday palms of subtropical South America, which retain old leaf sheaths as a persistent spiny coat around the stem. The ivory palms Phytelephas of northwestern South America produce seeds with a very hard white endosperm resembling ivory. The Ceroxyloideae are a primarily South American group with inflorescences accompanied by a protective prophyll and peduncular bracts.

Characters (from Healy & Edgar 1980): Usually arborescent, stout (and mostly unbranched) to slender, sometimes scandent and shrubby; usually monoecious, occasionally dioecious or bisexual. Leaves usually in terminal crowns, often very large, pinnate or palmate; leaflets narrow. Inflorescence usually paniculate, below or amongst leaves; spathes one or more, usually large. Flowers small, actinomorphic, more or less sessile, commonly unisexual, males and females sometimes in one inflorescence; sepals 3, free or connate; petals 3, free or connate. Stamens usually 6 in two whorls; anthers two-celled, dehiscing by longitudinal slits. Ovary superior, 1-3-celled or 3 carpels separate; ovule usually solitary. Fruit a berry or drupe.

<==Arecaceae [Areciflorae, Palmaceae, Palmae, Palmales, Principes]
    |--Trithrinax acanthocoma DS04
    |--+--Phytelephas aequatorialis DS04
    |  `--Arecoideae DS04
    `--+--Coryphoideae DS04
       |--Nypoideae [Nypaceae] CBH93
       |    |--Spinizonocolpites baculatus CBH93
       |    `--Nypa fruticans DS04
       |         |--N. f. var. fruticans H09
       |         `--N. f. var. neameana H09
       `--+--Plectocomia elongata DS04
          `--Calamoideae DS04

Arecaceae incertae sedis:
  Elaeis PJ02
    |--E. guineensis PJ02
    `--E. oleifera MS10
  Calyptrogyne L2001
  Metroxylon K03
    |--M. rumphii M72
    |--M. sagu K03 [incl. Sagus genuina L1801]
    `--M. salmonense Z93
  Plectocomiopsis geminiflora K03
  Veitchia H03
  Pritchardia remota O94
  Pelagodoxeae TB08
    |--Sommieria TB08
    `--Pelagodoxa H03
  Manicaria saccifera H03, O49
  Orbygnya DS86
  Scheclea DS86
  Geonoma BBB03
    |--G. cuneata BBB03
    `--G. irena BBB03
  Hyospathe DS86
  Lepidocaryoideae CBH93
    |--Lepidocaryum DS86
    |--Monocolpites franciscoi Muller 1970 CBH93
    `--Dicolpopollis CBH93
         |--D. elegans Muller 1979 CBH93
         `--D. malesianus Muller 1979 CBH93
  Astrocaryum jauari DS86
  Maximiliana DS86
  Desmoncus DS86
  Mauritia flexuosa DS86
  Acanthococos YY22
  Acanthophoenix YY22
  Acrocomia F11
    |--A. aculeata F11
    |--A. mexicana G69
    `--A. totai F11
  Actinokentia YY22
  Adelodypsis YY22
  Aeris YY22
  Asterogyne YY22
  Barbosa YY22
  Barcella YY22
  Calyptronoma YY22
  Catoblastus YY22
  Cocops YY22
  Cyphokentia YY22
  Cyphophoenix YY22
  Didymosperma YY22
  Diplothemium YY22
  Gigliolia YY22
  Haplodypsis YY22
  Haplophloga YY22
  Howea B00
    |--H. belmoreana B00
    `--H. forsterana P82
  Hyophorbe lagenicaulis Z93
  Juania YY22
  Jubaea YY22
  Kentiopsis YY22
  Kunthia YY22
  Linospadix YY22
  Leopoldinia YY22
  Martinezia YY22
  Mischophloeus YY22
  Neodypsis decaryi Z93
  Nephrocarpus YY22
  Oncocalamus YY22
  Oreodoxa YY22
  Phloga [incl. Neophloga] Z03
    `--P. nodifera Z93
  Phlogella YY22
  Podococcus YY22
  Pseudophoenix sargentii OB11
  Ptychoraphis YY22
  Ptychandra YY22
  Sclerosperma YY22
  Trichodypsis YY22
  Vershaffeltia YY22
  Vonitra YY22
  Wallichia YY22
  Welfia YY22
  Ancistrophyllum YY22
  Ceratolobus YY22
  Coleococcus YY22
  Nannorhops YY22
  Cyclospathe YY22
  Eugeissona P88
    |--E. insignis P88
    |--E. tristis P88
    `--E. utilis K03
  Kinetostigma YY22
  Lodoicea seychellarum YY22, C55
  Pholidocarpus YY22
  Ravenia YY22
  Solfia YY22
  Wendlandiella YY22
  Acanthorhiza YY22
  Rhapidophyllum YY22
  Rhapis YY22
  Trachycarpus fortunei (Hooker) Wendland 1862 HE80 [incl. Chamoerops excelsa (l. c.) LO98]
  Zalacca YY22
  Sabal BBB03
    |--S. bermudana WM14
    |--S. megacarpa BTA75
    |--S. minor I-BH-O10
    |--S. palmetto BBB03
    `--S. texana MG56
  Roystonea SK02
    |--R. borinquena SWK87
    |--R. hispaniolana HSS13
    |--R. princeps (Becc.) Burret 1929 CD07
    `--R. regia SK02
  Oraniopsis appendiculata E99
  Rhopalostylis Wendl. & Drude 1875 [=Eora Cook 1927] ME70
    |--R. baueri Wendl. & Drude 1875 ME70 [=Areca baueri Hook.f. 1868 C06, Kentia baueri C06]
    |--R. cheesemanii Becc. 1917 ME70
    `--R. sapida Wendl. & Drude in Kerch. 1878 ME70 [=Areca sapida C06, Kentia sapida C06]
  Ceroxyloideae HB09
    |--Synechanthus fibrosus Z03
    `--Ceroxylon [Ceroxyleae] HB09
         `--C. quindiuense HSS13
  Coccothrinax montana J87
  Washingtonia M93
    |--W. filifera M93
    `--W. robusta K10
  Pigafetta filaris M82
  Inodes mexicana G69
  Sabalites Saporta 1865 HL08
    `--‘Flabellaria’ major Unger 1847 (see below for synonymy) HL08
  Phoenicites Brongniart 1828 HL08
  Musophyllum axonense Watelet 1866 HL08
  Chrysalidocarpus Z93
    |--C. lutescens Z93
    `--C. madagascariensis Z93
  Aphanes caryotifolia Z93
  Carpentaria acuminata Z93
  Wodyetia bifurcata Z93
  Hedyscepe canterburyana B00
  Butia yatay F11
  Scheelea F11
    |--S. cephalotes F11
    `--S. princeps F11
  Eolirion primigenium Schenk 1869 CBH93
  Pilophora Jacq. 1802 KC01

‘Flabellaria’ major Unger 1847 [=Sabal major (Unger) Heer 1855; incl. S. (Flabellaria) haeringiana] HL08

*Type species of generic name indicated

REFERENCES

[BTA75] Baker, E. W., D. M. Tuttle & M. J. Abbatiello. 1975. The false spider mites of northwestern and north central Mexico (Acarina: Tenuipalpidae). Smithsonian Contributions to Zoology 194: 1–23.

[BBB03] Barfod, A. S., T. Burholt & F. Borchsenius. 2003. Contrasting pollination modes in three species of Licuala (Arecaceae: Coryphoideae). Telopea 10 (1): 207–223.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[CD07] Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1–E44.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[CBH93] Collinson, M. E., M. C. Boulter & P. L. Holmes. 1993. Magnoliophyta (‘Angiospermae’). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 809–841. Chapman & Hall: London.

[DS04] Davis, J. I., D. W. Stevenson, G. Petersen, O. Seberg, L. M. Campbell, J. V. Freudenstein, D. H. Goldman, C. R. Hardy, F. A. Michelangeli, M. P. Simmons, C. D. Specht, F. Vergara-Silva & M. Gandolfo. 2004. A phylogeny of the monocots, as inferred from rbcL and atpA sequence variation, and a comparison of methods for calculating jackknife and bootstrap values. Systematic Botany 29 (3): 467–510.

[DS86] Dixon, J. R. & P. Soini. 1986. The Reptiles of the Upper Amazon Basin, Iquitos Region, Peru. Milwaukee Public Museum: Milwaukee.

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.

[F11] Fraga, R. M. 2011. Family Icteridae (New World blackbirds). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 684–807. Lynx Edicions: Barcelona.

[G69] Goodwin, G. G. 1969. Mammals from the State of Oaxaca, Mexico, in the American Museum of Natural History. Bulletin of the American Museum of Natural History 141 (1): 1–269, 40 pls.

[H03] Heads, M. 2003. Ericaceae in Malesia: Vicariance biogeography, terrane tectonics and ecology. Telopea 10 (1): 311–449.

[HE80] Healy, A. J., & E. Edgar. 1980. Flora of New Zealand vol. 3. Adventive cyperaceous, petalous and spathaceous monocotyledons. P. D. Hasselberg, Government Printer: Wellington (New Zealand).

[HB09] Heatubun, C. D., W. J. Baker, J. P. Mogea, M. M. Harley, S. S. Tjitrosoedirdjo & J. Dransfield. 2009. A monograph of Cyrtostachys (Arecaceae). Kew Bulletin 64 (1): 67–94.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[HL08] Henniger, M. & R. Leder. 2008. Eozäne Makroflora der Querfurter Mulde. Mauritiana 20 (2): 229–251.

[HSS13] Hirschfeld, E., A. Swash & R. Still. 2013. The World's Rarest Birds. Princeton University Press: Princeton (New Jersey).

[I-BH-O10] Ibáñez-Bernal, S., & V. Hernández-Ortiz. 2010. Ropalomeridae (ropalomerid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1025–1030. NRC Research Press: Ottawa.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[K10] Keighery, G. 2010. The naturalised vascular plants of the Pilbara region, Western Australia. Records of the Western Australian Museum Supplement 78 (1): 299–311.

[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby's Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).

[K03] Kulip, J. 2003. An ethnobotanical survey of medicinal and other useful plants of Muruts in Sabah, Malaysia. Telopea 10 (1): 81–98.

[L1801] Labillardiere, C. 1801. Mémoire sur le sagoutier des Moluques et le cocotier des Maldives. Bulletin des Sciences, par la Societé Philomathique de Paris 2 (46): 170–171.

[LO98] Lack, H. W., & H. Ohba. 1998. Die Xylothek des Chikusai Kato. Willdenowia 28: 263–276.

[L2001] Lindquist, E. E. 2001. Poising for a new century: diversification in acarology. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 17–34. CSIRO Publishing: Melbourne.

[M72] McKean, J. L. 1972. Notes on some collections of bats (order Chiroptera) from Papua-New Guinea and Bougainville Island. CSIRO Division of Wildlife Research Technical Paper 26: 1–35.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[MG56] Mockford, E. L. & A. B. Gurney. 1956. A review of the psocids, or book-lice and bark-lice, of Texas (Psocoptera). Journal of the Washington Academy of Sciences 46 (11): 353–368.

[ME70] Moore, L. B., & E. Edgar. 1970. Flora of New Zealand vol. 2. Indigenous Tracheophyta: Monocotyledones except Gramineae. A. R. Shearer, Government Printer: Wellington (New Zealand).

[MS10] Moore, M. J., P. S. Soltis, C. D. Bell, J. G. Burleigh & D. E. Soltis. 2010. Phylogenetic analysis of 83 plastid genes further resolves the early diversification of eudicots. Proceedings of the National Academy of Sciences of the USA 107 (10): 4623–4628.

[M82] Musser, G. G. 1982. Results of the Archbold Expeditions. No. 110. Crunomys and the small-bodied shrew rats native to the Philippine Islands and Sulawesi (Celebes). Bulletin of the American Museum of Natural History 174 (1): 1–95.

[O49] Oliver, A. P. 1949. The fatty acid composition of Hemideina thoracica White. Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 321–324.

[OB11] Orenstein, R. I., & D. Brewer. 2011. Family Cardinalidae (cardinals). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 330–427. Lynx Edicions: Barcelona.

[O94] Otte, D. 1994. The Crickets of Hawaii: origin, systematics and evolution. The Orthopterists' Society: The Academy of Natural Sciences of Philadelphia.

[P82] Pickard, J. 1982. Catastrophic disturbance and vegetation on Little Slope, Lord Howe Island. Australian Journal of Ecology 7: 161–170.

[PJ02] Poinar, G. O., Jr., T. A. Jackson, N. L. Bell & M. B. Wahid. 2002. Elaeolenchus parthenonema n. g., n. sp. (Nematoda: Sphaerularioidea: Anandranematidae n. fam.) parasitic in the palm-pollinating weevil Elaeidobius kamerunincus Faust, with a phylogenetic synopsis of the Sphaerularioidea Lubbock 1861. Systematic Parasitology 52: 219–225.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[SWK87] Snyder, N. F. R., J. W. Wiley & C. B. Kepler. 1987. The Parrots of Luquillo: Natural history and conservation of the Puerto Rican parrot. Western Foundation of Vertebrate Zoology: Los Angeles.

[SK02] Sumithramma, N., A. R. V. Kumar, K. Chandrashekara & D. Rajagopal. 2002. Plant selection for nesting by Oecophylla smaragdina, Hymenoptera: Formicidae: do physical features affect the choice of the plant? Journal of the Bombay Natural History Society 99 (3): 408–412.

[TB08] Trudgen, M. S., & W. J. Baker. 2008. A revision of the Heterospathe elegans (Arecaceae) complex in New Guinea. Kew Bulletin 63 (4): 639–647.

[WM14] Wickett, N. H., S. Mirarab, N. Nguyen, T. Warnow, E. Carpenter, N. Matasci, S. Ayyampalayam, M. S. Barker, J. G. Burleigh, M. A. Gitzendanner, B. R. Ruhfel, E. Wafula, J. P. Der, S. W. Graham, S. Mathews, M. Melkonian, D. E. Soltis, P. S. Soltis, N. W. Miles, C. J. Rothfels, L. Pokorny, A. J. Shaw, L. DeGeronimo, D. W. Stevenson, B. Surek, J. C. Villarreal, B. Roure, H. Philippe, C. W. dePamphilis, T. Chen, M. K. Deyholos, R. S. Baucom, T. M. Kutchan, M. M. Augustin, J. Wang, Y. Zhang, Z. Tian, Z. Yan, X. Wu, X. Sun, G. K.-S. Wong & J. Leebens-Mack. 2014. Phylotranscriptomic analysis of the origin and early diversification of land plants. Proceedings of the National Academy of Sciences of the USA 111 (45): E4859–E4868.

[YY22] Yampolsky, C., & H. Yampolsky. 1922. Distribution of sex forms in the phanerogamic flora. Bibliotheca Genetica 3: 1–62.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

[Z03] Zona, S. 2003. Endosperm condition and the paradox of Ptychococcus paradoxus. Telopea 10 (1): 179–185.

Last updated: 21 December 2019.

Arecoideae

Mountain cabbage palm Prestoea montana, photographed by Jose D. Alicea.


Belongs within: Arecaceae.

The Arecoideae is the largest of the generally recognised palm subfamilies, containing about 1100 species. Notable members include the coconut Cocos nucifera, widely grown in tropical regions of the world for their large edible seeds, as well as for fibres from the leaves and fruit.

Synapomorphies (from the Angiosperm Phylogeny Website): Hypodermal cells hexagonal, transversely elongate; flowers in triads or in two vertical rows; style branches separate; n = 16.

<==Arecoideae
    |--Chamaedorea [Chamaedoreeae] HB09
    |    |--C. carchensis HB09
    |    `--C. microcarpa MH98
    |--Iriarteeae HB09
    |    |--Iriartea exerrhiza HB09, DS86
    |    `--Dictyocaryum HB09
    |--Euterpeae HB09
    |    |--Euterpe Z03
    |    |    |--E. edulis DS86
    |    |    `--E. oleracea DS04
    |    `--Prestoea HB09
    |         |--P. longepetiolata Z03
    |         |    |--P. l. var. longepetiolata Z03
    |         |    `--P. l. var. cuatrecasasii Z03
    |         |--P. montana [incl. Euterpe globosa] J87
    |         `--P. pubens Z03
    |--Cocoseae HB09
    |    |  i. s.: Cocos B88
    |    |           |--C. nucifera K03
    |    |           `--C. yattae B88
    |    |--Bactridinae HB09
    |    |    |--Bactris Z93
    |    |    |    |--B. gasipaes Z93
    |    |    |    `--B. utilis C38
    |    |    `--Aiphanes eggersii HB09, BBB03
    |    `--Attaleinae HB09
    |         |--Voanioala HB09
    |         `--Attalea HB09
    |              |--A. butyracea F11
    |              |--A. cohune G69
    |              |--A. maripa B14
    |              `--A. phalerata HSS13
    `--Areceae HB09
         |--Clinostigma HB09
         |--Nenga HB09
         |--Iguanura HB09
         |--Areca catechu HB09
         |--Hydriastele HB09
         |    |--H. costata HB09
         |    `--H. wendlandiana B00a
         |--Heterospathe [incl. Barkerwebbia Beccari 1905] TB08
         |    |--H. compsoclada (Burret) Heatubun in Heatubun, Baker et al. 2009 (see below for synonymy) HB09
         |    |--H. elegans (Beccari) Beccari 1909 (see below for synonymy) TB08
         |    |    |--H. e. ssp. elegans [incl. H. versteegiana Beccari 1909] TB08
         |    |    `--H. e. ssp. humilis (Beccari) Trudgen & Baker 2008 (see below for synonymy) TB08
         |    |--H. lepidota Moore 1969 TB08
         |    `--H. pullenii Trudgen & Baker 2008 TB08
         `--Cyrtostachys Blume 1838 [Cyrtostachydinae] HB09
              |  i. s.: C. ledermanniana Beccari 1923 HB09
              |--C. glauca Moore 1966 HB09
              `--+--*C. renda Blume 1838 (see below for synonymy) HB09
                 |--C. bakeri Heatubun in Heatubun, Baker et al. 2009 HB09
                 |--C. barbata Heatubun in Heatubun, Baker et al. 2009 HB09
                 |--C. elegans Burret 1937 HB09
                 |--C. excelsa Heatubun in Heatubun, Baker et al. 2009 HB09
                 `--C. loriae Beccari 1905 (see below for synonymy) HB09

Arecoideae incertae sedis:
  Wettinia Z03
  Reinhardtia Z03
  Dypsis Z03
  Neonicholsonia Z03
  Archontophoenix Z03
    |--A. alexandrae E99
    `--A. cunninghamiana B00a
  Laccospadix australasica Z03, B00a
  Calyptrocalyx Z03
  Drymophloeus [incl. Coleospadix] Z03
  Normanbya normabyi Z03, Z93
  Adonidia Z03
  Ptychosperma Z03
    |--P. bleeseri B00a
    |--P. elegans [incl. P. capitis-yorkii] Z03
    `--P. macarthurii B00a
  Loxococcus Z03
  Lemurophoenix Z03
  Siphokentia Z03
  Gulubia Z03
  Pinanga rumphiana Z03, HB09
  Rhopaloblaste Z03
    |--R. ceramica (Miq.) Burret 1928 [=Cyrtostachys ceramica (Miq.) Wendl. in Kerch. 1878] HB09
    `--R. ledermanniana HB09
  Dictyosperma album Z03, Z93
  Actinorhytis Z03
  Physokentia Z03
  Oncosperma tigillarium Z03, K03
  Verschaffeltia Z03
  Roscheria Z03
  Phoenicophorium Z03
  Nephrospermum Z03
  Beccariophoenix Z03
  Lytocaryum Z03
  Polyandrococos Z03
  Syagrus [incl. Rhyticocos] Z03
    |--S. coronata B04
    `--S. romanzoffiana Z93
  Oenocarpus [incl. Jessenia] Z03
    |--‘Jessenia’ batana DS86
    `--O. mapora SDK-L03
  Ptychococcus Z03
    |--P. arecinus [=Ptychosperma arecina Beccari 1877] Z03
    |--P. lepidotus Z03
    `--P. paradoxus (see below for synonymy) Z03
  Caryota Z03
    |--C. albertii B00b
    |--C. mitis P88
    |--C. no P88
    `--C. obtusifolia P88
         |--C. o. var. obtusifolia P88
         `--C. o. var. aequatorialis P88

Cyrtostachys loriae Beccari 1905 [incl. C. brassii Burret 1935, C. kisu Beccari 1914, C. microcarpa Burret 1939, C. peekeliana Beccari 1914, C. phanerolepis Burret 1936] HB09

*Cyrtostachys renda Blume 1838 [=Bentinckia renda (Blume) Mart. 1853; incl. Ptychosperma coccinea Teijsm. & Binn. 1866 (n. n.), Areca erythrocarpa Wendl. in Kerch. 1878 (n. n.), A. erythropoda Miq. 1861, Cyrtostachys lakka Becc. 1885, Pinanga purpurea Miq. 1861 (nom. inv.), P. rubricaulis Linden 1885 (n. n.), Cyrtostachys lakka var. singaporensis Becc. 1885] HB09

Heterospathe compsoclada (Burret) Heatubun in Heatubun, Baker et al. 2009 [=Cyrtostachys compsoclada Burret 1936] HB09

Heterospathe elegans (Beccari) Beccari 1909 [=*Barkerwebbia elegans Beccari 1905, B. elongata (l. c.), Heterospathe elongata (l. c.)] TB08

Heterospathe elegans ssp. humilis (Beccari) Trudgen & Baker 2008 [=H. humilis Beccari 1914, Barkerwebbia humilis (Beccari) Beccari ex Martelli 1935] TB08

Ptychococcus paradoxus [=Drymophloeus paradoxus Scheffer 1876, Ptychosperma paradoxa (Scheffer) Scheffer 1876] Z03

*Type species of generic name indicated

REFERENCES

[BBB03] Barfod, A. S., T. Burholt & F. Borchsenius. 2003. Contrasting pollination modes in three species of Licuala (Arecaceae: Coryphoideae). Telopea 10 (1): 207–223.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B00a] Braby, M. F. 2000a. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[B00b] Braby, M. F. 2000b. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).

[B04] Brown, B. V. 2004. Revision of the subgenus Udamochiras of Melaloncha bee-killing flies (Diptera: Phoridae: Metopininae). Zoological Journal of the Linnean Society 140: 1–42.

[C38] Crawford, J. C. 1938. A new genus and species of Thysanoptera from greenhouses. Proceedings of the Entomological Society of Washington 40 (4): 109–111.

[DS04] Davis, J. I., D. W. Stevenson, G. Petersen, O. Seberg, L. M. Campbell, J. V. Freudenstein, D. H. Goldman, C. R. Hardy, F. A. Michelangeli, M. P. Simmons, C. D. Specht, F. Vergara-Silva & M. Gandolfo. 2004. A phylogeny of the monocots, as inferred from rbcL and atpA sequence variation, and a comparison of methods for calculating jackknife and bootstrap values. Systematic Botany 29 (3): 467–510.

[DS86] Dixon, J. R. & P. Soini. 1986. The Reptiles of the Upper Amazon Basin, Iquitos Region, Peru. Milwaukee Public Museum: Milwaukee.

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.

[F11] Fraga, R. M. 2011. Family Icteridae (New World blackbirds). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 684–807. Lynx Edicions: Barcelona.

[G69] Goodwin, G. G. 1969. Mammals from the State of Oaxaca, Mexico, in the American Museum of Natural History. Bulletin of the American Museum of Natural History 141 (1): 1–269, 40 pls.

[HB09] Heatubun, C. D., W. J. Baker, J. P. Mogea, M. M. Harley, S. S. Tjitrosoedirdjo & J. Dransfield. 2009. A monograph of Cyrtostachys (Arecaceae). Kew Bulletin 64 (1): 67–94.

[HSS13] Hirschfeld, E., A. Swash & R. Still. 2013. The World's Rarest Birds. Princeton University Press: Princeton (New Jersey).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[K03] Kulip, J. 2003. An ethnobotanical survey of medicinal and other useful plants of Muruts in Sabah, Malaysia. Telopea 10 (1): 81–98.

[MH98] Morikawa, H., A. Higaki, M. Nohno, M. Takahashi, M. Kamada, M. Nakata, G. Toyohara, Y. Okamura, K. Matsui, S. Kitani, K. Fujita, K. Irifune & N. Goshima. 1998. More than a 600-fold variation in nitrogen dioxide assimilation among 217 plant taxa. Plant, Cell and Environment 21: 180–190.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[SDK-L03] Saw, L. G., J. Dransfield & D. M. Keith-Lucas. 2003. Morphological diversity of the genus Licuala (Palmae). Telopea 10 (1): 187–206.

[TB08] Trudgen, M. S., & W. J. Baker. 2008. A revision of the Heterospathe elegans (Arecaceae) complex in New Guinea. Kew Bulletin 63 (4): 639–647.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

[Z03] Zona, S. 2003. Endosperm condition and the paradox of Ptychococcus paradoxus. Telopea 10 (1): 179–185.

Last updated: 13 November 2018.